Updated on 2022/04/26
Doctor of Agriculture （ 1992.3 University of Tsukuba ）
Forest Genetics
Population Genetics
Molecular Ecology
Life Science / Forest science
Environmental Science/Agriculture Science / Conservation of biological resources
Population genomics of forest tree species
Breeding system of tree populations
Conservation genetics in forest tree species
Japan Society for Promotion of Science Research Center for Science Systems Researcher
2018.4 - 2022.3
Country：Japan
Nagoya University Graduate School of Bioagricultural Sciences Field Science Center Director in General
2013.4 - 2019.3
北海道立林業試験場 客員研究員
2007.9 - 2007.12
Country：Japan
Nagoya University Graduate School of Bioagricultural Sciences Professor
2005.4
Country：Japan
総合地球環境学研究所 共同研究員
2005.4 - 2011.3
Country：Japan
岐阜県森林科学研究所 客員研究員
2001.6 - 2002.3
Country：Japan
Nagoya University Graduate School of Bioagricultural Sciences Assistant Professor
1999.4 - 2005.3
Country：Japan
Nagoya University School of Agricultural Sciences Assistant Professor
1997.10 - 1999.3
Country：Japan
科学技術振興事業団 科学技術特別研究員
1995.10 - 1997.9
Country：Japan
University of Tsukuba Institute of Agriculture and Forestry Assistant
1992.8 - 1995.9
Country：Japan
University of Tsukuba Graduate School, Division of Agriculture
- 1992.3
Country： Japan
University of Tsukuba Second Cluster of College College of Agriculture and Forestry
- 1987.3
Country： Japan
The Japanese Forest Society
2016.4
Japanese Society of Forest Genetics and Tree Breeding President
2018.4 - 2020.3
Ecological Society of Japan
Tree Genetics & Genomes Associate Editor
2019.12
Botanical Society of America
Botanical Society of Japan
Society for Study of Species Biology
Japanese Association of Historical Botany
中部森林学会 会長
2021.6
林木育種賞
2009.5 社団法人林木育種協会
5S-IGS rDNA in wind-pollinated trees (Fagus L.) encapsulates 55 million years of reticulate evolution and hybrid origins of modern species Reviewed
Cardoni Simone, Piredda Roberta, Denk Thomas, Grimm Guido W., Papageorgiou Aristotelis C., Schulze Ernst-Detlef, Scoppola Anna, Shanjani Parvin Salehi, Suyama Yoshihisa, Tomaru Nobuhiro, Worth James R. P., Simeone Marco Cosimo
PLANT JOURNAL Vol. 109 ( 4 ) page： 909 - 926 2022.2
Determination of intraspecific variation in seed weight, leaf functional traits, and sapling size of Betula ermanii using a common garden experiment Reviewed
Paing Aye Myat Myat, Chen Shufen, Tsumura Yoshihiko, Tomaru Nobuhiro, Homma Kousuke, Kadomatsu Masahiko, Yoshida Toshiya, Kobayashi Hajime, Iio Atsuhiro, Osumi Katsuhiro, Taneda Haruhiko, Hisamoto Yoko, Goto Susumu
JOURNAL OF FOREST RESEARCH Vol. 26 ( 6 ) page： 419 - 426 2021.11
Reciprocal crosses between Magnolia stellata and Magnolia kobus do not show significant reproductive barriers in seed formation Reviewed
Tamaki Ichiro, Wadasaki Naotaka, Ishida Kiyoshi, Tomaru Nobuhiro
PLANT SPECIES BIOLOGY Vol. 36 ( 4 ) page： 596 - 601 2021.10
Assessments of fine-scale spatial patterns of SNPs in an old-growth beech forest. Reviewed
Tukamoto, M., Akada, S., Matsuda, S., Jouyu, H., Kisanuki, H., Tomaru, N., and Torimaru, T.
Heredity Vol. 125 page： 240–252 2020.6
Hotspots of stream tadpole diversity in forest and agricultural landscapes in Ranomafana, Madagascar Reviewed
Ramamonjisoa Noelikanto, Sakai Masaru, Ndriantsoa Serge Herilala, Kakehashi Ryosuke, Kurabayashi Atsushi, Tomaru Nobuhiro, Natuhara Yosihiro
LANDSCAPE AND ECOLOGICAL ENGINEERING Vol. 16 page： 207–221 2020.1
Spatial variation in bird pollination and its mitigating effects on the genetic diversity of pollen pools accepted by Camellia japonica trees within a population at a landscape level.
Nakanishi A, Takeuchi T, Ueno S, Nishimura N, Tomaru N
Heredity Vol. 124 page： 170-181 2020.1
The origin and genetic variability of vegetatively propagated clones identified from old planted trees and plantations of Thujopsis dolabrata var. hondae in Ishikawa Prefecture, Japan
Ikeda Torazo, Mishima Kentaro, Takata Katsuhiko, Tomaru Nobuhiro
TREE GENETICS & GENOMES Vol. 15 page： 80 2019.12
Population genetic structure and demography of Magnolia kobus: variety borealis is not supported genetically
Tamaki Ichiro, Kawashima Naomichi, Setsuko Suzuki, Lee Jung-Hyun, Itaya Akemi, Yukitoshi Kyohei, Tomaru Nobuhiro
JOURNAL OF PLANT RESEARCH Vol. 132 ( 6 ) page： 741-758 2019.11
The complete chloroplast genome of Fagus crenata (subgenus Fagus) and comparison with F. engleriana (subgenus Engleriana)
Worth James R. P., Liu Luxian, Wei Fu-Jin, Tomaru Nobuhiro
PEERJ Vol. 7 page： e7026 2019.6
Development of microsatellite markers from genomic DNA of Parashorea malaanonan (Diptero-carpaceae) using next-generation sequencing
Tinio Crusty E., Ueno Saneyoshi, Uchiyama Kentaro, San-Jose Maldia Lerma, Tomaru Nobuhiro
SILVAE GENETICA Vol. 68 ( 1 ) page： 22-25 2019.1
Morphological and genetic divergence between two lineages of Magnolia salicifolia (Magnoliaceae) in Japan Reviewed
Tamaki Ichiro, Kawashima Naomichi, Setsuko Suzuki, Itaya Akemi, Tomaru Nobuhiro
BIOLOGICAL JOURNAL OF THE LINNEAN SOCIETY Vol. 125 ( 3 ) page： 475-490 2018.11
Identification of Thujopsis dolabrata var. hondae clones and their distribution across plantations in Ishikawa prefecture Reviewed
Ikeda Torazo, Tokuda Noriko, Tomaru Nobuhiro
JOURNAL OF FOREST RESEARCH Vol. 23 ( 2 ) page： 105-111 2018
Reduced incompatibility in the production of second generation hybrids between two Magnolia species revealed by Bayesian gene dispersal modeling Reviewed
Tamaki, I., Tani, S., Setsuko, S., Ueno, S., Wadasaki, N., and Tomaru, N.
American Journal of Botany Vol. 104 page： 1546-1555 2017
Development and characterization of EST-SSR markers for the genus Rhododendron section Brachycalyx (Ericaceae) Reviewed
Yoichi, W., Sakaguchi, S., Ueno, S., Tomaru, N., and Uehara, K.
Plant Species Biology Vol. 32 page： 455-459 2017
Contrasting diversification history between insular and continental species of three-leaved azaleas (Rhododendron sect. Brachycalyx) in East Asia Reviewed
Yoichi, W., Jin, X.-F., Peng, C.-I., Tamaki, I., and Tomaru, N.
Journal of Biogeography Vol. 44 page： 065-1076 2017
Sex change in the subdioecious shrub Eurya japonica (Pentaphylacaceae) Reviewed
Wang, H., Matsushita, M., Tomaru N., and Nakagawa, M.
Ecology and Evolution Vol. 7 page： 2340-2345 2017
Genetic diversity and structure of remnant Magnolia stellata populations affected by anthropogenic pressures and a conservation strategy for maintaining their current genetic diversity Reviewed
Tamaki, I., Setsuko, S., and Tomaru, N.
Conservation Genetics Vol. 17 page： 715-725 2016
Thinning operations increase the demographic performance of the rare subtree species Magnolia stellata in a suburban forest landscape Reviewed
Matsushita, M., Setsuko, S., Tamaki, I., Nakagawa, M., Nishimura, N., and Tomaru, N.
Landscape and Ecological Engineering Vol. 12 page： 179-186 2016
Pollen dispersal patterns and population persistence in a small isolated population of Fagus crenata Reviewed
Inanaga. M., Koyama, Y., Ida, H., Okada, M., Nakanishi, A., Takahashi, M., and Tomaru, N.
Tree Genomes & Genetics Vol. 12 page： 69 2016
High male fertility in males of a subdioecious shrub in hand-pollinated crosses Reviewed
Wang, H., Matsushita, M., Tomaru N., and Nakagawa, M.
AoB Plants Vol. 8 page： plw067 2016
Population demographic history of a temperate shrub, Rhododendron weyrichii (Ericaceae), on continental islands of Japan and South Korea Reviewed
Yoichi, W., Tamaki, I., Sakaguchi, S., Song, Yamamoto, J., and Tomaru, N.
Ecology and Evolution Vol. 6 page： 8800-8810 2016
Forest dynamics of stands damaged by Japanese oak wilt in the Kaisho Forest, Aichi Prefecture Reviewed
Watanabe, N., Okada, T., Tomaru, N., Nishimura, N., and Nakagawa, M.
Journal of the Japanese Forest Society Vol. 98 page： 273-278 2016
Family-specific responses in survivorship and phenotypic traits to different light environments in a seedling population of Fagus crenata in a cool-temperate forest Reviewed
Torimaru, T., Takeda, Y., Matsushita, M., Tamaki, I., Sano, J., and Tomaru, N.
Population Ecology Vol. 57 page： 77-91 2015
Differences in female reproductive success between female and hermaphrodite individuals in the subdioecious shrub Eurya japonica (Theaceae) Reviewed
Wang, H., Matsushita, M., Tomaru N., and Nakagawa M.
Plant Biology Vol. 17 page： 194-200 2015
Inbreeding depression at the sapling stage and its genetic consequences in a population of the outcrossing dominant tree species, Castanopsis sieboldii. Reviewed
Nakanishi, A., Yoshimaru, H., Tomaru, N., Miura, M., Manabe, T., and Yamamoto S.
Tree Genetics & Genomes Vol. 11 page： 62 2015
Distance-dependent but genetically random mating in a Japanese beech (Fagus crenata) population Reviewed
Inanaga, M., Nakanishi, A., Torimaru, T., Nishimura, N., and Tomaru, N.
Botany Vol. 92 page： 795-803 2014
Conflict in outcomes for conservation based on population genetic diversity and genetic divergence approaches: a case study in the Japanese relictual conifer Sciadopitys verticillata (Sciadopityaceae) Reviewed
Worth, J. R. P., Yokogawa, M., Pérez-Figueroa, A., Tsumura, Y., Tomaru, N., Janes, J. K., and Isagi, Y.
Conservation Genetics Vol. 15 page： 1243-1257 2014
Differences in seed formation and germination rates between reciprocal interspecific crosses in Magnolia stellata and M. salicifolia Reviewed
Tani, S., Tamaki, I., Setsuko, S., and Tomaru, N.
Journal of the Japanese Forest Society Vol. 96 page： 200-205 2014
Genetic lineage and diversity of Fagus crenata trees planted in Shizuoka Prefecture Reviewed
Katai, H., Yamada, S., Hiraoka, K., Hoshikawa, T., Tomaru, N., and Takahashi, M.
Forest Genetics and Tree Breeding Vol. 3 page： 101-110 2014
Patterns of geographic distribution have a considerable influence on population genetic structure in one common and two rare species of Rhododendron (Ericaceae) Reviewed
Yoichi, W. and Tomaru, N.
Tree Genetics & Genomes Vol. 10 page： 827-837 2014
Indigenous genetic lineages of Fagus crenata found in the Izu Peninsula suggest that there was one of refugia for the species during the last glacial maximum Reviewed
Katai, H., Takahashi, M., Hiraoka, K., Yamada, Y., and Tomaru, N.
Journal of Forest Research Vol. 18 page： 418-429 2013
Asymmetric introgression between Magnolia stellata and M. salicifolia at a site where the two species grow sympatrically Reviewed
Muranishi, S., Tamaki, I., Setsuko, S., and Tomaru, N.
Tree Genetics & Genomes Vol. 9 page： 1005-1015 2013
Pollen flow and effects of population structure on selfing rates and female and male reproductive success in fragmented Magnolia stellata populations Reviewed
Setsuko, S., Nagamitsu, T., and Tomaru, N.
BMC Ecology Vol. 13 page： 10 2013
Genetic diversity and population genetic structure in widespread deciduous broad-leaved trees in Japan Invited Reviewed
Tomaru, N.
Chikyu Kankyo Vol. 18 page： 119-126 2013
Japanese beech (Fagus crenata) plantations established from seedlings of non-native genetic lineages Reviewed
Koyama, Y., Takahashi, M., Murauchi, Y., Fukatsu, E., Watanabe, A., and Tomaru, N.
Journal of Forest Research Vol. 17 ( 1 ) page： 116-120 2012
Reproductive investment at stem and genet levels in male and female plants of the clonal dioecious shrub Ilex leucoclada (Aquifoliaceae) Reviewed
Torimaru, T. and Tomaru, N.
Botany Vol. 90 ( 4 ) page： 301-310 2012
Effects of pollen availability on pollen immigration and pollen donor diversity in riparian dioecious trees (Salix arbutifolia) Reviewed
Hoshikawa, T., Nagamitsu, T., and Tomaru, N.
Botany Vol. 90 ( 6 ) page： 481-489 2012
Patterns of pollen flow in a dense population of the insect-pollinated canopy tree species Castanopsis sieboldii Reviewed
Nakanishi, A., Yoshimaru, H., Tomaru, N., Miura, M., Manabe, T., and Yamamoto S.
Journal of Heredity Vol. 103 ( 4 ) page： 547-556 2012
Differences in clonal integration between the sexes: long term demographic patterns in the dioecious, multi-stemmed shrub Lindera triloba Reviewed
Matsushita, M. and Tomaru, N.
Botany Vol. 90 ( 10 ) page： 1028-1035 2012
Sexual differences in year-to-year flowering trends in the dioecious multi-stemmed shrub Lindera triloba: effects of light and clonal integration Reviewed
Matsushita, M., Nakagawa, M., and Tomaru, N.
Journal of Ecology Vol. 99 ( 6 ) page： 1520–1530 2011
The effects of plant size and light availability on male and female reproductive success and functional gender in a hermaphrodite tree species, Magnolia stellata Reviewed
Setsuko, S. and Tomaru, N.
Botany Vol. 89 ( 9 ) page： 593-604 2011
Inference of genetic lineages of Fagus crenata populations in Shizuoka Prefecture based on chloroplast DNA and nuclear microsatellite variations Reviewed
Katai, H., Takahashi, M., Hiraoka, K., Yamada, S., Yamamoto, S., Kato, K., Hakamada, T., and Tomaru, N.
Journal of the Japanese Forest Society Vol. 93 ( 2 ) page： 73-78 2011
Factors affecting the production, growth and survival of sprouting stems in the multi-stemmed understory shrub, Lindera triloba Reviewed
Matsushita, M, Tomaru, N, Hoshino, D, Nishimura, N., and Yamamoto, S.
Botany Vol. 88 ( 2 ) page： 174-184 2010
Genetic structure of an endemic Japanese conifer, Sciadopitys verticillata (Sciadopityaceae), by using microsatellite markers Reviewed
Kawase, D., Tsumura, Y., Tomaru, N., Seo, A., and Yumoto, T.
Journal of Heredity Vol. 101 ( 3 ) page： 292-297 2010
*Genetic divergence in nuclear genomes between populations of Fagus crenata along the Japan Sea and Pacific sides of Japan Reviewed
Hiraoka, K. and Tomaru, N.
Journal of Plant Research Vol. 122 ( 3 ) page： 269-282 2009
Development and characterization of EST-SSR markers for Sciadopitys verticillata (Sciadopityaceae) Reviewed
Kawase, D., Ueno, S., Tsumura, T., Tomaru, N., Seo, A., and Yumoto, T.
Conservation Genetics Vol. 10 ( 6 ) page： 1997-1999 2009
Mitochondrial DNA variation in natural populations of Japanese larch (Larix kaempferi) Reviewed
San Jose-Maldia, L., Uchida, K., and Tomaru, N.
Silvae Genetica Vol. 58 ( 5/6 ) page： 234-241 2009
Development of EST-SSR markers from an inner bark cDNA library of Fagus crenata (Fagaceae). Reviewed
Ueno, S. Taguchi, Y., Tomaru, N., and Tsumura, Y.
Conservation Genetics Vol. 10 ( 5 ) page： 1477-1485 2009
Population genetic structure of Fagus japonica revealed by nuclear microsatellite markers Reviewed
Hiraoka, K. and Tomaru, N.
International Journal of Plant Sciences Vol. 170 ( 6 ) page： 748-758 2009
Eighteen microsatellite loci in Salix arbutifolia (Salicaceae) and cross-species amplification in Salix and Populus species Reviewed
Hoshikawa, T, Kikuchi, S., Nagamitsu, T., and Tomaru, N.
Molecular Ecology Resources Vol. 9 ( 4 ) page： 1202-1205 2009
*Effects of seed- and pollen-mediated gene dispersal on genetic structure among Quercus salicina saplings Reviewed
Nakanishi, A., Tomaru, N., Yoshimaru, H., Manabe, T., and Yamamoto S.
Heredity Vol. 102 ( 2 ) page： 182-189 2009
Estimation of outcrossing rates at hierarchical levels of fruits, individuals, populations and species in Magnolia stellata Reviewed
Tamaki, I., Setsuko, S. and Tomaru, N.
Heredity Vol. 102 ( 4 ) page： 381-388 2009
*Inter-population variation in mating system and late-stage inbreeding depression in Magnolia stellata Reviewed
Tamaki, I., Ishida, K., Setsuko, S., and Tomaru, N.
Molecular Ecology Vol. 18 ( 6 ) page： 2365-2374 2009
Relationships between flowering phenology and female reproductive success in the Japanese tree species Magnolia stellata Reviewed
Setsuko, S., Tamaki, I., Ishida, K., and Tomaru, N.
Botany Vol. 86 ( 3 ) page： 248-258 2008
*Genetic variation and differentiation in populations of a threatened tree, Magnolia stellata: factors influencing the level of within-population genetic variation Reviewed
Tamaki, I., Setsuko, S., and Tomaru, N.
Heredity Vol. 100 ( 4 ) page： 415-423 2008
The soil seed bank of the threatened plant Magnolia stellata is subordinate to emergence of current-year seedlings Reviewed
Kisanuki, H., Oguro, H., Nakai, A., Setsuko, S., Nishimura, N., and Tomaru, N.
Journal of Forest Research Vol. 13 ( 2 ) page： 143-146 2008
Population differentiation and gene flow within a metapopulation of a threatened tree, Magnolia stellata (Magnoliaceae). Reviewed
Setsuko, S., Ishida, K., Ueno, S., Tsumura Y., and Tomaru, N.
American Journal of Botany Vol. 94 ( 1 ) page： 128-136 2007
*Effects of kin-structured seed dispersal on the genetic structure of the clonal shrub Ilex leucoclada Reviewed
Torimaru, T., Tani, N., Tsumura, Y. Nishimura, N., and Tomaru, N.
Evolution Vol. 61 ( 6 ) page： 1289-1300 2007
Pollen-mediated gene flow in a small, fragmented natural population of Fagus crenata Blume. Reviewed
Hanaoka, S., Yuzurihara, J., Asuka, Y., Tomaru, N., Tsumura, Y. Kakubari, T., and Mukai, Y.
Canadian Journal of Botany Vol. 85 ( 4 ) page： 404-413 2007
Reduced seed production, inbreeding, and pollen shortage in a small population of a threatened tree, Magnolia stellata Reviewed
Hirayama, K. Ishida, K., Setsuko, S., and Tomaru, N.
Biological Conservation Vol. 136 ( 2 ) page： 315-323 2007
Effects of canopy gaps on the genetic structure of Camellia japonica saplings in a Japanese old-growth evergreen forest Reviewed
Ueno, S., Tomaru, N., Yoshimaru, H., Manabe, T., and Yamamoto, S.
Heredity Vol. 96 ( 4 ) page： 304-310 2006
Relationships between flowering phenology, plant size and female reproductive output in a dioecious shrub, Ilex leucoclada (Aquifoliaceae) Reviewed
Torimaru, T. and Tomaru, N.
Canadian Journal of Botany Vol. 84 ( 12 ) page： 1860-1869 2006
Development and polymorphisms of microsatellite markers for hinoki (Chamaecyparis obtusa) Reviewed
Matsumoto, A., Tani, N., Li, X-G, Nakao, Y., Tomaru, N., and Tsumura, Y.
Molecular Ecology Notes Vol. 6 ( 2 ) page： 310-312 2006
Fine-scale clonal structure and diversity within patches of a clone-forming dioecious shrub, Ilex leucoclada (Aquifoliaceae) Reviewed
Torimaru, T. and Tomaru, N.
Annals of Botany Vol. 95 ( 2 ) page： 295-304 2005
Interannual genetic heterogeneity of pollen pools accepted by Quercus salicina individuals Reviewed
Nakanishi, A., Tomaru, N., Yoshimaru, H., Manabe, T., and Yamamoto S.
Molecular Ecology Vol. 14 ( 15 ) page： 4469-4478 2005
Half-sib family structure of Fagus crenata saplings in an old-growth beech-dwarf bamboo forest Reviewed
Asuka, Y, Tomaru, N., Munehara, Y., Tani, N., Tsumura, Y., and Yamamoto S.
Molecular Ecology Vol. 14 ( 8 ) page： 2565-2575 2005
Development of microsatellite markers in Magnolia stellata. (Magnoliaceae), a threatened Japanese tree Reviewed
Setsuko, S., Ueno, S., Tsumura Y., and Tomaru, N.
Conservation Genetics Vol. 6 ( 2 ) page： 317-320 2005
Effects of pollen shortage and self-pollination on seed production of an endangered tree, Magnolia stellata Reviewed
Hirayama, K. Ishida, K., and Tomaru, N.
Annals of Botany Vol. 95 ( 6 ) page： 1009-1015 2005
Development and characterization of microsatellite markers for Fagus crenata Blume Reviewed
Asuka, Y., Tani, N., Tsumura, Y., and Tomaru, N.
Molecular Ecology Notes Vol. 4 ( 1 ) page： 101-103 2004
Patterns of pollen flow and genetic differentiation among pollen pools in Quercus salicina in a warm temperate old-growth evergreen broad-leaved forest Reviewed
Nakanishi, A., Tomaru, N., Yoshimaru, H., Kawahara, T., Manabe, T. and Yamamoto, S
Silvae Genetica Vol. 53 ( 5/6 ) page： 258-264 2004
Development and polymorphism of simple sequence repeat DNA markers for the evergreen shrub Ilex leucoclada M. Reviewed
Torimaru, T., Tani, N., Tsumura, Y., Hiraoka K., and Tomaru, N.
Molecular Ecology Notes Vol. 4 ( 3 ) page： 531-533 2004
Size distribution and genetic structure in relation to clonal growth within a population of Magnolia tomentosa Thunb. (Magnoliaceae) Reviewed
Setsuko, S., Ishida, K., and Tomaru, N.
Molecular Ecology Vol. 13 ( 9 ) page： 2645-2653 2004
Stem distribution and clonal structure of Chamaecyparis pisifera growing in an old-growth beech-conifer forest Reviewed
Hayakawa, T., Tomaru, N., and Yamamoto, S.
Ecological Research Vol. 19 ( 4 ) page： 411-420 2004
Heterogeneous genetic structure in a Fagus crenata population in an old-growth beech forest revealed by microsatellite markers Reviewed
Asuka, Y., Tomaru, N., Nishimura, N., Tsumura, Y., and Yamamoto, S.
Molecular Ecology Vol. 13 ( 5 ) page： 1241-1250 2004
Comparison of the fine-scale genetic structure of three dipterocarp species Reviewed
Takeuchi, Y., Ichikawa, S., Konuma, A., Tomaru, T., Niiyama, K., Lee, S. L., Muhammad, N., and Tsumura, Y.
Heredity Vol. 92 ( 4 ) page： 323-328 2004
Clonal diversity and genetic differentiation in Ilex leucoclada M. pathces in an old-growth beech forest Reviewed
Torimaru, T., Tomaru, N., Nishimura N., and Yamamoto S.
Molecular Ecology Vol. 12 ( 4 ) page： 809-818 2003
Genetic diversity of nuclear and mitochondrial genomes in Pinus parviflora Sieb. & Zucc. (Pinaceae) populations Reviewed
Tani, N., Maruyama, K., Tomaru, N., Uchida, K., Araki, M., Tsumura, Y., Yoshimaru, Y., and Ohba, K.
Heredity Vol. 91 ( 11 ) page： 510-518 2003
Genetic structure in the northernmost marginal population of Japanese beech (Fagus crenata Blume): Influence of the founding event on genetic structure Reviewed
Takahashi, M., Tomaru, N., Ubukata, M., and Koono, K.
Silvae Genetica Vol. 51 ( 5-6 ) page： 219-225 2002
Size-class differences in genetic structure and individual distribution of Camellia japonica L. in a Japanese old-growth evergreen forest Reviewed
Ueno, S., Tomaru, N., Yoshimaru, H., Manabe, T., and Yamamoto, S.
Heredity Vol. 89 ( 2 ) page： 120-126 2002
Chloroplast DNA phylogeography of Fagus crenata (Fagaceae) in Japan Reviewed
Fujii N., Tomaru, N., Okuyama, K., Koike, T, Mikami, T., and Ueda, K.
Plant Systematics and Evolution Vol. 232 ( 1-2 ) page： 21-33 2002
Highy polymorphic microsatellite markers in Chamaecyparis obtusa Reviewed
Nakao, U., Iwata, H., Matsumoto, Y., Tsumura, Y., and Tomaru, N
Canadian Journal of Forest Research Vol. 31 ( 12 ) page： 2248-2251 2001
Genetic structure of Camellia japonica L. in an old-growth evergreen forest, Tsushima, Japan Reviewed
Ueno, S., Tomaru, N., Yoshimaru, H., Manabe, T., and Yamamoto, S.
Molecular Ecology Vol. 9 ( 7 ) page： 647-656 2000
Genetic diversity and differentiation of Taxodium in the south-eastern United States using cleaved amplified polymorphic sequences Reviewed
Tsumura, Y., Tomaru, T., Suyama, Y., and Bacchus, S.
Heredity Vol. 83 ( 3 ) page： 229-238 1999
Development and characterization of microsatellite markers in Camellia japonica L. Reviewed
Ueno S, Yoshimaru H, Tomaru N, and Yamamoto S.
Molecular Ecology Vol. 8 ( 2 ) page： 335-336 1999
Genetic diversity of Cryptomeria japonica using co-dominant DNA markers based on Sequenced-Tagged Site Reviewed
Tsumura, Y. and Tomaru, N.
Theoretical and Applied Genetics Vol. 98 ( 3-4 ) page： 396-404 1999
Genetic structure within a Japanese stone pine (Pinus pumila Regel) population on Mt. Aino-dake in central Honshu, Japan Reviewed
Tani, N., Tomaru, N., Tsumura, Y., Araki, M., and Ohba, K
Journal of Plant Research Vol. 111 ( 1101 ) page： 7-15 1998
Detection of quantitative trait loci for juvenile growth, flower bearing and rooting ability based on a linkage map in sugi Reviewed
Yoshimaru, H, Ohba, K., Tsurumi, K., Tomaru, N., Murai, M, Mukai, Y., Suyama, Y., Tsumura. Y., Kawahara, T., and Sakamaki, Y.
Theoretical and Applied Genetics Vol. 97 ( 1-2 ) page： 45-50 1998
Intraspecific variation and phylogeographic patterns of Fagus crenata (Fagaceae) mitochorial DNA Reviewed
Tomaru, N., Takahashi, M., Tsumura, Y., Takahashi, M., and Ohba, K
American Journal of Botany Vol. 85 ( 5 ) page： 629-636 1998
Genetic diversity in Fagus crenata (Japanese beech): influence of the distributional shift during the late-Quaternary Reviewed
Tomaru, N., Mitsutsuji, T., Takahashi, M., Tsumura, Y., Uchida, K., and Ohba, K.
Heredity Vol. 78 ( 3 ) page： 241-251 1997
Linkage analysis of isozyme and dwarf loci, and detection of lethal genes in sugi (Cryptomeria japonica D. Don) Reviewed
Kuramoto, N., Tomaru, N., Murai, M., and Ohba, K.
Breeding Science Vol. 47 ( 3 ) page： 259-266 1997
Allozyme variation in natural populations of hinoki, Chamaecyparis obtusa and its comparison with hinoki plus-trees selected from artificial stands Reviewed
Uchida, K., Tomaru, N., Tomaru, T., Yamamoto, C., and Ohba, K.
Breeding Science Vol. 47 ( 1 ) page： 7-14 1997
Regeneration system and genetic diversity of Cryptomeria japonica at different growing altitudes Reviewed
Taira, H., Tsumura, Y., Tomaru, N., and Ohba, K.
Canadian Journal of Forest Research Vol. 27 ( 4 ) page： 447-452 1997
Genetic diversity and differentiation in populations of Japanese stone pine (Pinus pumila) in Japan Reviewed
Tani, N., Tomaru, N., Araki, M., and Ohba, K.
Canadian Journal of Forest Research Vol. 26 ( 8 ) page： 1454-1462 1996
Genetic analysis of four isozyme loci and a linkage relationship between dwarf gene (dw) and leaf whiting gene in summer (swl) of sugi (Cryptomeria japonica) Reviewed
Kuramoto, N., Tomaru, N., Murai, M., Ohba, K.
Breeding Science Vol. 46 ( 4 ) page： 379-384 1996
The genetic variation of isozyme in natural populations of masson pine. Reviewed
Huang, Q., Wang, L., Tomaru, N., and Ohba, K.
Acta Genetica Sinica Vol. 22 ( 2 ) page： 142-151 1995
Molecular phylogeny of conifers using RFLP analysis of PCR-amplified specific chloroplast genes Reviewed
Tsumura, Y., Yoshimura, K., Tomaru, N., and Ohba, K.
Theoretical and Applied Genetics Vol. 91 ( 8 ) page： 1222-1236 1995
Allozyme variations and spatial distributions of genotypes in strains of a butt-rot fungus of Japanese larch Reviewed
Kuroda, Y., Tomaru, N., and Hanyu, M.
Journal of the Japanese Forestry Society Vol. 77 ( 5 ) page： 480-485 1995
A linkage map for sugi(Cryptomeria japonica)based on RFLP, RAPD and isozyme loci Reviewed
Mukai, Y., Suyama, Y., Tsumura, Y., Kawahara, T., Yoshimaru, H., Kondo, T., Tomaru, N., Kuramoto, N., and Murai, M.
Theoretical and Applied Genetics Vol. 90 ( 6 ) page： 835-840 1995
Correlations between the growth vigour and isozyme phenotypes of Pinus massoniana Lamb. Reviewed
Huang, Q., Wang, L., Tomaru, N. and Ohba, K.
Scientia Silvae Sinicae Vol. 30 ( 3 ) page： 214-219 1994
Genetic control of isozyme variation in masson pine, Pinus massoniana Lamb Reviewed
Huang, Q.-Q., Tomaru. N., Wang, L.-H., and Ohba, K.
Silvae Genetica Vol. 43 ( 5-6 ) page： 285-292 1994
Genetic variation and population differentiation in natural populations of Cryptomeria japonica Reviewed
Tomaru, N., Tsumura, Y., and Ohba, K.
Plant Species Biology Vol. 9 ( 3 ) page： 191-199 1994
Studies on genetic variation in plantations and plus-trees of Cryptomeria Japonica using isozyme markers Reviewed
Tomaru, N.
Memoirs of The Institue of Agriculture and Forestry, University of Tsukuba Vol. 6 page： 1-66 1994
Allozyme variation in plus-tree of hinoki, Chamaecyparis obtusa, selected from artificial stands Reviewed
Uchida, K., Tomaru, N., Tsumura, Y., Takahashi, C. and Ohba, K.
Japanese Journal of Breeding Vol. 43 ( 4 ) page： 485-494 1993
Allozyme variation in artificial stands and a plus-tree group of sugi, Cryptomeria japonica in Ibaraki Prefecture Reviewed
Tomaru, N., Tsumura, Y. and Ohba, K.
Journal of the Japanese Forestry Society Vol. 74 ( 1 ) page： 44-48 1992
Inheritance of isozyme variants in Korean pine(Pinus koraiensis) Reviewed
Tomaru, N., Tsumura, Y. and Ohba, K.
Journal of the Japanese Forestry Society Vol. 72 ( 3 ) page： 194-200 1990
森林学の百科事典. 日本森林学会編
戸丸信弘（ Role： Contributor）
丸善出版 2021.1
地図でわかる樹木の種苗移動ガイドライン. 津村義彦・陶山佳久編
戸丸信弘（ Role： Contributor , カラマツ、シデコブシ、ブナ、イヌブナ）
文一総合出版 2015.6
森林樹木の遺伝的変異と地史的分布変遷. 井出雄二・大河内勇・井上真編 教養として森林学
戸丸信弘（ Role： Sole author）
文永堂出版 2014
種内の遺伝的変異−中立変異−. 井出雄二・白石進編 森林遺伝育種学
戸丸信弘, 津村義彦（ Role： Joint author）
文永堂出版 2012
種内の遺伝的変異 –適応的変異−. 井出雄二・白石進編 森林遺伝育種学
津村義彦, 戸丸信弘（ Role： Joint author）
文永堂出版 2012
Conservation of Biological and Genetic Resources by Conservation Genetic Approaches (Considering Biodiversity as Resources. Imura, H. ed.) (in Japanese)
Nobuhiro Tomaru（ Role： Sole author）
Cokusai Shoin Co., Ltd. 2010
Genetic Variation and Structure in Fagus crenata populations (Applied Ecology for Restoration for Beech Forests. Terazawa, K. and Koyama, H. eds.) (in Japanese)
Nobuhiro Tomaru（ Role： Sole author）
Bun-ichi Sogo Shuppan Co. 2008
Encyclopedia of Plant Breeding (Japanese Society of Breeding ed.) (in Japanese)
Nobuhiro Tomaru（ Role： Contributor）
Baifukan Co., Ltd. 2005
Fagus crenata, Cryptomeria japonica, and Chamaecyparis obtusa (Tree Ecophysiology Koike, T. ed.) (in Japanese)
Nobuhiro Tomaru（ Role： Sole author）
Asakura Publishing Co., Ltd. 2004
Forest management and conservation using microsatellite markers: the example of Fagus (Biotechnology in Agriculture and Forestry, Vol. 55 Molecular Marker Systems. Lorz, H. L. and Wenzel, G. eds.)
Tsumura, Y., Takahashi, M., Takahashi, T., Tani, N., Asuka Y. and Tomaru N.（ Role： Joint author）
Springer-Verlag 2004
Encyclopedia of Ecology (Iwasa, Y., Matsumoto, T., Kikuzawa, K., and Ecological Society of Japan eds.) (in Japanese)
Nobuhiro Tomaru（ Role： Contributor）
Kyoritsu Shuppan, Co., Ltd. 2003
History of genes: History and Genetic Variation of Fagus crenata populations (Molecular Ecology of Woody Species. The Society for the Study of Species Biology ed.) (in Japanese)
Nobuhiro Tomaru（ Role： Sole author）
Bun-ichi Sogo Shuppan Co. 2001
広葉樹における遺伝的多様性の評価手法の開発
2010.4 - 2012.3
森林環境保全総合対策事業
Grant type：Competitive
自然再生事業のための遺伝的多様性の評価技術を用いた植物の遺伝的ガイドラインに関する研究
2005.4 - 2010.3
環境省地球環境保全等試験研究
Grant type：Competitive
人為的要因によって小集団化した希少樹種の保全管理技術に関する研究
2005.4 - 2008.3
環境省地球環境保全等試験研究
Grant type：Competitive
絶滅が危惧される希少樹種の生息域内保全に関する基礎的研究
2002.4 - 2005.3
環境省地球環境保全等試験研究
Grant type：Competitive
Grant number：21H04732 2021.4 - 2024.3
Grants-in-Aid for Scientific Research Grant-in-Aid for Scientific Research (A)
Authorship：Coinvestigator(s)
Improving climate change impact prediction by considering genetic variability within species
Grant number：21K05643 2021.4 - 2024.3
Grants-in-Aid for Scientific Research Grant-in-Aid for Scientific Research (C)
Authorship：Coinvestigator(s)
環境適応遺伝変異の空間モデリングによる主要高木種11種の将来気候下でのリスク評価
Grant number：21H02240 2021.4 - 2024.3
科学研究費助成事業 基盤研究(B)
内山 憲太郎, 津村 義彦, 戸丸 信弘, 加藤 珠理
Authorship：Coinvestigator(s)
今後数十年の間に予想される急速な温暖化に対する森林樹木の脆弱性の評価は、森林生態系保全上の喫緊の課題である。そこで本研究では、日本全国に広く分布する森林の主要構成樹種11種を対象に、気候変動に対する樹木の応答を、環境適応に関わる遺伝変異と気候データの空間モデリングを通して明らかにする。また、それらの解析を通して、それぞれの樹種において、気候変動の影響を特に強く受ける地域を地図上で明らかにし、今後の森林管理の指針に資する情報を提供する。
Elucidation of adaptive and neutral evolution processes in Fagus crenata using population genomics
Grant number：20H03027 2020.4 - 2023.3
Grant-in-Aid for Scientific Research Grant-in-Aid for Scientific Research(B)
Authorship：Principal investigator
国内外来種コブシが希少種シデコブシの存続に及ぼす影響
2017.4 - 2020.3
科学研究費補助金 基盤研究(B)
戸丸信弘
Authorship：Principal investigator
ブナ林の断片化がブナ集団の遺伝的多様性と繁殖に及ぼす影響
2013.4 - 2016.3
科学研究費補助金 基盤研究(B)
戸丸信弘
Authorship：Principal investigator
ブナ集団における適応的遺伝子の探索と生態的意義
2008.4 - 2011.3
科学研究費補助金 基盤研究(B)
戸丸 信弘
Authorship：Principal investigator
葉緑体ゲノムのSNPを用いたブナの分子系統地理学的・環境適応的研究
2007.4 - 2010.3
科学研究費補助金 基盤研究（B)
高橋誠
Authorship：Coinvestigator(s)
針葉樹の雑種苗の分子識別と起源推定
2007.4 - 2010.3
科学研究費補助金 基盤研究(B)
津村義彦
Authorship：Coinvestigator(s)
フタバガキ科の系統地理学的研究と産地識別のための塩基配列データベースの構築
2006.4 - 2010.3
科学研究費補助金 基盤研究（A)（海外学術調査）
津村義彦
Authorship：Coinvestigator(s)
湿地林を構成する希少木本種の繁殖と更新に及ぼす遺伝的荷重の影響の解明
2006.4 - 2009.3
科学研究費補助金 基盤研究（B)
石田清
Authorship：Coinvestigator(s)
絶滅危惧種シデコブシの保全を目指した遺伝子流動と近交弱勢に関する研究
2004.4 - 2007.3
科学研究費補助金 基盤研究(B)
戸丸 信弘
Authorship：Principal investigator
森林生態系の維持機構に関する個体群統計学・分子集団遺伝学的研究
2003.4 - 2004.3
科学研究費補助金 基盤研究（C)(1)（企画調査）
山本進一
Authorship：Coinvestigator(s)
分子生態遺伝学的手法による森林動態に関する研究
2002.4 - 2005.3
科学研究費補助金 基盤研究(A)
山本進一
Authorship：Coinvestigator(s)
メソスケールでの森林現象動態モニタリング技術の基盤確立
2001.4 - 2004.3
科学研究費補助金 基盤研究(A)(2)(展開)
山本進一
Authorship：Coinvestigator(s)
日本海側ブナ林構成樹種における地理的分布と遺伝的構造の歴史的関連性
2000.4 - 2003.3
科学研究費補助金 基盤研究(C)(2)(一般)
戸丸信弘
Authorship：Principal investigator
マイクロサテライトマーカーによる照葉樹天然林の繁殖構造解析
1999.4 - 2002.3
科学研究費補助金 基盤研究(B)(2)(一般)
山本進一
Authorship：Coinvestigator(s)
オルガネラDNA解析によるブナ集団の系統地理学的研究
1998.4 - 2000.3
科学研究費補助金 基盤研究（C)
戸丸信弘
Authorship：Principal investigator
環境問題と人間
2021
Conservation Genetics
2021
Forest Resource Management 1
2021
Plant Taxonomy and Vegetation Science
2021
Laboratory and Practice in Bioenvironmental Sciences 1
2021
Laboratory and Practice in Bioenvironmental Sciences 2
2021
Forest Biology
2021
Basic Practical Courses
2021
Environmental Problems and Human Beings
2020
Conservation Genetics
2020
Forest Resource Management 1
2020
Laboratory and Practice in Bioenvironmental Sciences 1
2020
Laboratory and Practice in Bioenvironmental Sciences 2
2020
Forest Biology
2020
Laboratory/Field Course in Forest Biology
2020
森林資源環境学特論
2014.4 - 2015.3 （Mie University）
特別講義
2000.4 - 2001.3 （Ehime University）
特別講義
1999.4 - 2000.3 （Niigata University）
