岐阜県森林科学研究所   客員研究員

2001.6 - 2002.3

Nagoya University   Graduate School of Bioagricultural Sciences   Associate Professor

1999.4 - 2005.3

Nagoya University   School of Agricultural Sciences   Associate Professor

1997.10 - 1999.3

科学技術振興事業団   科学技術特別研究員

1995.10 - 1997.9

University of Tsukuba   Institute of Agriculture and Forestry   Assistant

1992.8 - 1995.9

Botanical Society of Japan

Society for Study of Species Biology

Japanese Association of Historical Botany

中部森林学会   会長

2021.6 - 2024.3

5S-IGS rDNA in wind-pollinated trees (<i>Fagus</i> L.) encapsulates 55 million years of reticulate evolution and hybrid origins of modern species Reviewed

Cardoni, S; Piredda, R; Denk, T; Grimm, GW; Papageorgiou, AC; Schulze, ED; Scoppola, A; Shanjani, PS; Suyama, Y; Tomaru, N; Worth, JRP; Simeone, MC

PLANT JOURNAL   Vol. 109 ( 4 ) page： 909 - 926   2022.2

Determination of intraspecific variation in seed weight, leaf functional traits, and sapling size of <i>Betula ermanii</i> using a common garden experiment Reviewed

Paing, AMM; Chen, SF; Tsumura, Y; Tomaru, N; Homma, K; Kadomatsu, M; Yoshida, T; Kobayashi, H; Iio, A; Osumi, K; Taneda, H; Hisamoto, Y; Goto, S

JOURNAL OF FOREST RESEARCH   Vol. 26 ( 6 ) page： 419 - 426   2021.11

Reciprocal crosses between <i>Magnolia stellata</i> and <i>Magnolia kobus</i> do not show significant reproductive barriers in seed formation Reviewed

Tamaki, I; Wadasaki, N; Ishida, K; Tomaru, N

PLANT SPECIES BIOLOGY   Vol. 36 ( 4 ) page： 596 - 601   2021.10

Assessments of fine-scale spatial patterns of SNPs in an old-growth beech forest. Reviewed

Tukamoto, M., Akada, S., Matsuda, S., Jouyu, H., Kisanuki, H., Tomaru, N., and Torimaru, T.

Heredity   Vol. 125 ( 4 ) page： 240 - 252   2020.10

Hotspots of stream tadpole diversity in forest and agricultural landscapes in Ranomafana, Madagascar Reviewed

Ramamonjisoa Noelikanto, Sakai Masaru, Ndriantsoa Serge Herilala, Kakehashi Ryosuke, Kurabayashi Atsushi, Tomaru Nobuhiro, Natuhara Yosihiro

LANDSCAPE AND ECOLOGICAL ENGINEERING   Vol. 16 ( 3 ) page： 207 - 221   2020.7

Spatial variation in bird pollination and its mitigating effects on the genetic diversity of pollen pools accepted by Camellia japonica trees within a population at a landscape level. Reviewed

Nakanishi A, Takeuchi T, Ueno S, Nishimura N, Tomaru N

Heredity   Vol. 124 ( 1 ) page： 170 - 181   2020.1

The origin and genetic variability of vegetatively propagated clones identified from old planted trees and plantations of Thujopsis dolabrata var. hondae in Ishikawa Prefecture, Japan Reviewed

Ikeda Torazo, Mishima Kentaro, Takata Katsuhiko, Tomaru Nobuhiro

TREE GENETICS & GENOMES   Vol. 15 ( 6 ) page： 80   2019.12

Population genetic structure and demography of Magnolia kobus: variety borealis is not supported genetically Reviewed

Tamaki Ichiro, Kawashima Naomichi, Setsuko Suzuki, Lee Jung-Hyun, Itaya Akemi, Yukitoshi Kyohei, Tomaru Nobuhiro

JOURNAL OF PLANT RESEARCH   Vol. 132 ( 6 ) page： 741 - 758   2019.11

The complete chloroplast genome of Fagus crenata (subgenus Fagus) and comparison with F. engleriana (subgenus Engleriana) Reviewed

Worth James R. P., Liu Luxian, Wei Fu-Jin, Tomaru Nobuhiro

PEERJ   Vol. 7   page： e7026   2019.6

Development of microsatellite markers from genomic DNA of Parashorea malaanonan (Diptero-carpaceae) using next-generation sequencing Reviewed

Tinio Crusty E., Ueno Saneyoshi, Uchiyama Kentaro, San-Jose Maldia Lerma, Tomaru Nobuhiro

SILVAE GENETICA   Vol. 68 ( 1 ) page： 22-25   2019.1

Morphological and genetic divergence between two lineages of Magnolia salicifolia (Magnoliaceae) in Japan Reviewed

Tamaki Ichiro, Kawashima Naomichi, Setsuko Suzuki, Itaya Akemi, Tomaru Nobuhiro

BIOLOGICAL JOURNAL OF THE LINNEAN SOCIETY   Vol. 125 ( 3 ) page： 475-490   2018.11

Identification of Thujopsis dolabrata var. hondae clones and their distribution across plantations in Ishikawa prefecture Reviewed

Ikeda Torazo, Tokuda Noriko, Tomaru Nobuhiro

JOURNAL OF FOREST RESEARCH   Vol. 23 ( 2 ) page： 105-111   2018

Development and characterization of EST-SSR markers for the genus Rhododendron section Brachycalyx (Ericaceae) Reviewed

Yoichi, W., Sakaguchi, S., Ueno, S., Tomaru, N., and Uehara, K.

Plant Species Biology   Vol. 32 ( 4 ) page： 455 - 459   2017.10

Reduced incompatibility in the production of second generation hybrids between two Magnolia species revealed by Bayesian gene dispersal modeling Reviewed

Tamaki, I., Tani, S., Setsuko, S., Ueno, S., Wadasaki, N., and Tomaru, N.

American Journal of Botany   Vol. 104 ( 10 ) page： 1546 - 1555   2017.10

Contrasting diversification history between insular and continental species of three-leaved azaleas (Rhododendron sect. Brachycalyx) in East Asia Reviewed

Yoichi, W., Jin, X.-F., Peng, C.-I., Tamaki, I., and Tomaru, N.

Journal of Biogeography   Vol. 44 ( 5 ) page： 1065 - 1076   2017.5

Sex change in the subdioecious shrub Eurya japonica (Pentaphylacaceae) Reviewed

Wang, H., Matsushita, M., Tomaru N., and Nakagawa, M.

Ecology and Evolution   Vol. 7 ( 7 ) page： 2340 - 2345   2017.4

Genetic diversity and structure of remnant Magnolia stellata populations affected by anthropogenic pressures and a conservation strategy for maintaining their current genetic diversity Reviewed

Tamaki, I., Setsuko, S., and Tomaru, N.

Conservation Genetics   Vol. 17   page： 715-725   2016

Thinning operations increase the demographic performance of the rare subtree species Magnolia stellata in a suburban forest landscape Reviewed

Matsushita, M., Setsuko, S., Tamaki, I., Nakagawa, M., Nishimura, N., and Tomaru, N.

Landscape and Ecological Engineering   Vol. 12   page： 179-186   2016

Pollen dispersal patterns and population persistence in a small isolated population of Fagus crenata Reviewed

Inanaga. M., Koyama, Y., Ida, H., Okada, M., Nakanishi, A., Takahashi, M., and Tomaru, N.

Tree Genomes & Genetics   Vol. 12   page： 69   2016

High male fertility in males of a subdioecious shrub in hand-pollinated crosses Reviewed

Wang, H., Matsushita, M., Tomaru N., and Nakagawa, M.

AoB Plants   Vol. 8   page： plw067   2016

Population demographic history of a temperate shrub, Rhododendron weyrichii (Ericaceae), on continental islands of Japan and South Korea Reviewed

Yoichi, W., Tamaki, I., Sakaguchi, S., Song, Yamamoto, J., and Tomaru, N.

Ecology and Evolution   Vol. 6   page： 8800-8810   2016

Forest dynamics of stands damaged by Japanese oak wilt in the Kaisho Forest, Aichi Prefecture Reviewed

Watanabe, N., Okada, T., Tomaru, N., Nishimura, N., and Nakagawa, M.

Journal of the Japanese Forest Society   Vol. 98   page： 273-278   2016

Family-specific responses in survivorship and phenotypic traits to different light environments in a seedling population of Fagus crenata in a cool-temperate forest Reviewed

Torimaru, T., Takeda, Y., Matsushita, M., Tamaki, I., Sano, J., and Tomaru, N.

Population Ecology   Vol. 57   page： 77-91   2015

Differences in female reproductive success between female and hermaphrodite individuals in the subdioecious shrub Eurya japonica (Theaceae) Reviewed

Wang, H., Matsushita, M., Tomaru N., and Nakagawa M.

Plant Biology   Vol. 17   page： 194-200   2015

Inbreeding depression at the sapling stage and its genetic consequences in a population of the outcrossing dominant tree species, Castanopsis sieboldii. Reviewed

Nakanishi, A., Yoshimaru, H., Tomaru, N., Miura, M., Manabe, T., and Yamamoto S.

Tree Genetics & Genomes   Vol. 11   page： 62   2015

Distance-dependent but genetically random mating in a Japanese beech (Fagus crenata) population Reviewed

Inanaga, M., Nakanishi, A., Torimaru, T., Nishimura, N., and Tomaru, N.

Botany   Vol. 92   page： 795-803   2014

Conflict in outcomes for conservation based on population genetic diversity and genetic divergence approaches: a case study in the Japanese relictual conifer Sciadopitys verticillata (Sciadopityaceae) Reviewed

Worth, J. R. P., Yokogawa, M., Pérez-Figueroa, A., Tsumura, Y., Tomaru, N., Janes, J. K., and Isagi, Y.

Conservation Genetics   Vol. 15   page： 1243-1257   2014

Differences in seed formation and germination rates between reciprocal interspecific crosses in Magnolia stellata and M. salicifolia Reviewed

Tani, S., Tamaki, I., Setsuko, S., and Tomaru, N.

Journal of the Japanese Forest Society   Vol. 96   page： 200-205   2014

Genetic lineage and diversity of Fagus crenata trees planted in Shizuoka Prefecture Reviewed

Katai, H., Yamada, S., Hiraoka, K., Hoshikawa, T., Tomaru, N., and Takahashi, M.

Forest Genetics and Tree Breeding   Vol. 3   page： 101-110   2014

Patterns of geographic distribution have a considerable influence on population genetic structure in one common and two rare species of Rhododendron (Ericaceae) Reviewed

Yoichi, W. and Tomaru, N.

Tree Genetics & Genomes   Vol. 10   page： 827-837   2014

Indigenous genetic lineages of Fagus crenata found in the Izu Peninsula suggest that there was one of refugia for the species during the last glacial maximum Reviewed

Katai, H., Takahashi, M., Hiraoka, K., Yamada, Y., and Tomaru, N.

Journal of Forest Research   Vol. 18   page： 418-429   2013

Asymmetric introgression between Magnolia stellata and M. salicifolia at a site where the two species grow sympatrically Reviewed

Muranishi, S., Tamaki, I., Setsuko, S., and Tomaru, N.

Tree Genetics & Genomes   Vol. 9   page： 1005-1015   2013

Pollen flow and effects of population structure on selfing rates and female and male reproductive success in fragmented Magnolia stellata populations Reviewed

Setsuko, S., Nagamitsu, T., and Tomaru, N.

BMC Ecology   Vol. 13   page： 10   2013

Genetic diversity and population genetic structure in widespread deciduous broad-leaved trees in Japan Invited Reviewed

Tomaru, N.

Chikyu Kankyo   Vol. 18   page： 119-126   2013

Japanese beech (Fagus crenata) plantations established from seedlings of non-native genetic lineages Reviewed

Koyama, Y., Takahashi, M., Murauchi, Y., Fukatsu, E., Watanabe, A., and Tomaru, N.

Journal of Forest Research   Vol. 17 ( 1 ) page： 116-120   2012

Reproductive investment at stem and genet levels in male and female plants of the clonal dioecious shrub Ilex leucoclada (Aquifoliaceae) Reviewed

Torimaru, T. and Tomaru, N.

Botany   Vol. 90 ( 4 ) page： 301-310   2012

Effects of pollen availability on pollen immigration and pollen donor diversity in riparian dioecious trees (Salix arbutifolia) Reviewed

Hoshikawa, T., Nagamitsu, T., and Tomaru, N.

Botany   Vol. 90 ( 6 ) page： 481-489   2012

Patterns of pollen flow in a dense population of the insect-pollinated canopy tree species Castanopsis sieboldii Reviewed

Nakanishi, A., Yoshimaru, H., Tomaru, N., Miura, M., Manabe, T., and Yamamoto S.

Journal of Heredity   Vol. 103 ( 4 ) page： 547-556   2012

Differences in clonal integration between the sexes: long term demographic patterns in the dioecious, multi-stemmed shrub Lindera triloba Reviewed

Matsushita, M. and Tomaru, N.

Botany   Vol. 90 ( 10 ) page： 1028-1035   2012

Inference of genetic lineages of Fagus crenata populations in Shizuoka Prefecture based on chloroplast DNA and nuclear microsatellite variations Reviewed

Katai, H., Takahashi, M., Hiraoka, K., Yamada, S., Yamamoto, S., Kato, K., Hakamada, T., and Tomaru, N.

Journal of the Japanese Forest Society   Vol. 93 ( 2 ) page： 73-78   2011

Sexual differences in year-to-year flowering trends in the dioecious multi-stemmed shrub Lindera triloba: effects of light and clonal integration Reviewed

Matsushita, M., Nakagawa, M., and Tomaru, N.

Journal of Ecology   Vol. 99 ( 6 ) page： 1520–1530   2011

The effects of plant size and light availability on male and female reproductive success and functional gender in a hermaphrodite tree species, Magnolia stellata Reviewed

Setsuko, S. and Tomaru, N.

Botany   Vol. 89 ( 9 ) page： 593-604   2011

Factors affecting the production, growth and survival of sprouting stems in the multi-stemmed understory shrub, Lindera triloba Reviewed

Matsushita, M, Tomaru, N, Hoshino, D, Nishimura, N., and Yamamoto, S.

Botany   Vol. 88 ( 2 ) page： 174-184   2010

Genetic structure of an endemic Japanese conifer, Sciadopitys verticillata (Sciadopityaceae), by using microsatellite markers Reviewed

Kawase, D., Tsumura, Y., Tomaru, N., Seo, A., and Yumoto, T.

Journal of Heredity   Vol. 101 ( 3 ) page： 292-297   2010

*Genetic divergence in nuclear genomes between populations of Fagus crenata along the Japan Sea and Pacific sides of Japan Reviewed

Hiraoka, K. and Tomaru, N.

Journal of Plant Research   Vol. 122 ( 3 ) page： 269-282   2009

Development and characterization of EST-SSR markers for Sciadopitys verticillata (Sciadopityaceae) Reviewed

Kawase, D., Ueno, S., Tsumura, T., Tomaru, N., Seo, A., and Yumoto, T.

Conservation Genetics   Vol. 10 ( 6 ) page： 1997-1999   2009

Mitochondrial DNA variation in natural populations of Japanese larch (Larix kaempferi) Reviewed

San Jose-Maldia, L., Uchida, K., and Tomaru, N.

Silvae Genetica   Vol. 58 ( 5/6 ) page： 234-241   2009

Development of EST-SSR markers from an inner bark cDNA library of Fagus crenata (Fagaceae). Reviewed

Ueno, S. Taguchi, Y., Tomaru, N., and Tsumura, Y.

Conservation Genetics   Vol. 10 ( 5 ) page： 1477-1485   2009

Population genetic structure of Fagus japonica revealed by nuclear microsatellite markers Reviewed

Hiraoka, K. and Tomaru, N.

International Journal of Plant Sciences   Vol. 170 ( 6 ) page： 748-758   2009

Eighteen microsatellite loci in Salix arbutifolia (Salicaceae) and cross-species amplification in Salix and Populus species Reviewed

Hoshikawa, T, Kikuchi, S., Nagamitsu, T., and Tomaru, N.

Molecular Ecology Resources   Vol. 9 ( 4 ) page： 1202-1205   2009

*Effects of seed- and pollen-mediated gene dispersal on genetic structure among Quercus salicina saplings Reviewed

Nakanishi, A., Tomaru, N., Yoshimaru, H., Manabe, T., and Yamamoto S.

Heredity   Vol. 102 ( 2 ) page： 182-189   2009

Estimation of outcrossing rates at hierarchical levels of fruits, individuals, populations and species in Magnolia stellata Reviewed

Tamaki, I., Setsuko, S. and Tomaru, N.

Heredity   Vol. 102 ( 4 ) page： 381-388   2009

*Inter-population variation in mating system and late-stage inbreeding depression in Magnolia stellata Reviewed

Tamaki, I., Ishida, K., Setsuko, S., and Tomaru, N.

Molecular Ecology   Vol. 18 ( 6 ) page： 2365-2374   2009

Relationships between flowering phenology and female reproductive success in the Japanese tree species Magnolia stellata Reviewed

Setsuko, S., Tamaki, I., Ishida, K., and Tomaru, N.

Botany   Vol. 86 ( 3 ) page： 248-258   2008

*Genetic variation and differentiation in populations of a threatened tree, Magnolia stellata: factors influencing the level of within-population genetic variation Reviewed

Tamaki, I., Setsuko, S., and Tomaru, N.

Heredity   Vol. 100 ( 4 ) page： 415-423   2008

The soil seed bank of the threatened plant Magnolia stellata is subordinate to emergence of current-year seedlings Reviewed

Kisanuki, H., Oguro, H., Nakai, A., Setsuko, S., Nishimura, N., and Tomaru, N.

Journal of Forest Research   Vol. 13 ( 2 ) page： 143-146   2008

Population differentiation and gene flow within a metapopulation of a threatened tree, Magnolia stellata (Magnoliaceae). Reviewed

Setsuko, S., Ishida, K., Ueno, S., Tsumura Y., and Tomaru, N.

American Journal of Botany   Vol. 94 ( 1 ) page： 128-136   2007

*Effects of kin-structured seed dispersal on the genetic structure of the clonal shrub Ilex leucoclada Reviewed

Torimaru, T., Tani, N., Tsumura, Y. Nishimura, N., and Tomaru, N.

Evolution   Vol. 61 ( 6 ) page： 1289-1300   2007

Pollen-mediated gene flow in a small, fragmented natural population of Fagus crenata Blume. Reviewed

Hanaoka, S., Yuzurihara, J., Asuka, Y., Tomaru, N., Tsumura, Y. Kakubari, T., and Mukai, Y.

Canadian Journal of Botany   Vol. 85 ( 4 ) page： 404-413   2007

Reduced seed production, inbreeding, and pollen shortage in a small population of a threatened tree, Magnolia stellata Reviewed

Hirayama, K. Ishida, K., Setsuko, S., and Tomaru, N.

Biological Conservation   Vol. 136 ( 2 ) page： 315-323   2007

Effects of canopy gaps on the genetic structure of Camellia japonica saplings in a Japanese old-growth evergreen forest Reviewed

Ueno, S., Tomaru, N., Yoshimaru, H., Manabe, T., and Yamamoto, S.

Heredity   Vol. 96 ( 4 ) page： 304-310   2006

Relationships between flowering phenology, plant size and female reproductive output in a dioecious shrub, Ilex leucoclada (Aquifoliaceae) Reviewed

Torimaru, T. and Tomaru, N.

Canadian Journal of Botany   Vol. 84 ( 12 ) page： 1860-1869   2006

Development and polymorphisms of microsatellite markers for hinoki (Chamaecyparis obtusa) Reviewed

Matsumoto, A., Tani, N., Li, X-G, Nakao, Y., Tomaru, N., and Tsumura, Y.

Molecular Ecology Notes   Vol. 6 ( 2 ) page： 310-312   2006

Fine-scale clonal structure and diversity within patches of a clone-forming dioecious shrub, Ilex leucoclada (Aquifoliaceae) Reviewed

Torimaru, T. and Tomaru, N.

Annals of Botany   Vol. 95 ( 2 ) page： 295-304   2005

Interannual genetic heterogeneity of pollen pools accepted by Quercus salicina individuals Reviewed

Nakanishi, A., Tomaru, N., Yoshimaru, H., Manabe, T., and Yamamoto S.

Molecular Ecology   Vol. 14 ( 15 ) page： 4469-4478   2005

Half-sib family structure of Fagus crenata saplings in an old-growth beech-dwarf bamboo forest Reviewed

Asuka, Y, Tomaru, N., Munehara, Y., Tani, N., Tsumura, Y., and Yamamoto S.

Molecular Ecology   Vol. 14 ( 8 ) page： 2565-2575   2005

Development of microsatellite markers in Magnolia stellata. (Magnoliaceae), a threatened Japanese tree Reviewed

Setsuko, S., Ueno, S., Tsumura Y., and Tomaru, N.

Conservation Genetics   Vol. 6 ( 2 ) page： 317-320   2005

Effects of pollen shortage and self-pollination on seed production of an endangered tree, Magnolia stellata Reviewed

Hirayama, K. Ishida, K., and Tomaru, N.

Annals of Botany   Vol. 95 ( 6 ) page： 1009-1015   2005

Development and characterization of microsatellite markers for Fagus crenata Blume Reviewed

Asuka, Y., Tani, N., Tsumura, Y., and Tomaru, N.

Molecular Ecology Notes   Vol. 4 ( 1 ) page： 101-103   2004

Patterns of pollen flow and genetic differentiation among pollen pools in Quercus salicina in a warm temperate old-growth evergreen broad-leaved forest Reviewed

Nakanishi, A., Tomaru, N., Yoshimaru, H., Kawahara, T., Manabe, T. and Yamamoto, S

Silvae Genetica   Vol. 53 ( 5/6 ) page： 258-264   2004

Development and polymorphism of simple sequence repeat DNA markers for the evergreen shrub Ilex leucoclada M. Reviewed

Torimaru, T., Tani, N., Tsumura, Y., Hiraoka K., and Tomaru, N.

Molecular Ecology Notes   Vol. 4 ( 3 ) page： 531-533   2004

Size distribution and genetic structure in relation to clonal growth within a population of Magnolia tomentosa Thunb. (Magnoliaceae) Reviewed

Setsuko, S., Ishida, K., and Tomaru, N.

Molecular Ecology   Vol. 13 ( 9 ) page： 2645-2653   2004

Stem distribution and clonal structure of Chamaecyparis pisifera growing in an old-growth beech-conifer forest Reviewed

Hayakawa, T., Tomaru, N., and Yamamoto, S.

Ecological Research   Vol. 19 ( 4 ) page： 411-420   2004

Heterogeneous genetic structure in a Fagus crenata population in an old-growth beech forest revealed by microsatellite markers Reviewed

Asuka, Y., Tomaru, N., Nishimura, N., Tsumura, Y., and Yamamoto, S.

Molecular Ecology   Vol. 13 ( 5 ) page： 1241-1250   2004

Comparison of the fine-scale genetic structure of three dipterocarp species Reviewed

Takeuchi, Y., Ichikawa, S., Konuma, A., Tomaru, T., Niiyama, K., Lee, S. L., Muhammad, N., and Tsumura, Y.

Heredity   Vol. 92 ( 4 ) page： 323-328   2004

Clonal diversity and genetic differentiation in Ilex leucoclada M. pathces in an old-growth beech forest Reviewed

Torimaru, T., Tomaru, N., Nishimura N., and Yamamoto S.

Molecular Ecology   Vol. 12 ( 4 ) page： 809-818   2003

Genetic diversity of nuclear and mitochondrial genomes in Pinus parviflora Sieb. & Zucc. (Pinaceae) populations Reviewed

Tani, N., Maruyama, K., Tomaru, N., Uchida, K., Araki, M., Tsumura, Y., Yoshimaru, Y., and Ohba, K.

Heredity   Vol. 91 ( 11 ) page： 510-518   2003

Genetic structure in the northernmost marginal population of Japanese beech (Fagus crenata Blume): Influence of the founding event on genetic structure Reviewed

Takahashi, M., Tomaru, N., Ubukata, M., and Koono, K.

Silvae Genetica   Vol. 51 ( 5-6 ) page： 219-225   2002

Size-class differences in genetic structure and individual distribution of Camellia japonica L. in a Japanese old-growth evergreen forest Reviewed

Ueno, S., Tomaru, N., Yoshimaru, H., Manabe, T., and Yamamoto, S.

Heredity   Vol. 89 ( 2 ) page： 120-126   2002

Chloroplast DNA phylogeography of Fagus crenata (Fagaceae) in Japan Reviewed

Fujii N., Tomaru, N., Okuyama, K., Koike, T, Mikami, T., and Ueda, K.

Plant Systematics and Evolution   Vol. 232 ( 1-2 ) page： 21-33   2002

Highy polymorphic microsatellite markers in Chamaecyparis obtusa Reviewed

Nakao, U., Iwata, H., Matsumoto, Y., Tsumura, Y., and Tomaru, N

Canadian Journal of Forest Research   Vol. 31 ( 12 ) page： 2248-2251   2001

Genetic structure of Camellia japonica L. in an old-growth evergreen forest, Tsushima, Japan Reviewed

Ueno, S., Tomaru, N., Yoshimaru, H., Manabe, T., and Yamamoto, S.

Molecular Ecology   Vol. 9 ( 7 ) page： 647-656   2000

Genetic diversity and differentiation of Taxodium in the south-eastern United States using cleaved amplified polymorphic sequences Reviewed

Tsumura, Y., Tomaru, T., Suyama, Y., and Bacchus, S.

Heredity   Vol. 83 ( 3 ) page： 229-238   1999

Genetic diversity of Cryptomeria japonica using co-dominant DNA markers based on Sequenced-Tagged Site Reviewed

Tsumura, Y. and Tomaru, N.

Theoretical and Applied Genetics   Vol. 98 ( 3-4 ) page： 396-404   1999

Development and characterization of microsatellite markers in Camellia japonica L. Reviewed

Ueno S, Yoshimaru H, Tomaru N, and Yamamoto S.

Molecular Ecology   Vol. 8 ( 2 ) page： 335-336   1999

Genetic structure within a Japanese stone pine (Pinus pumila Regel) population on Mt. Aino-dake in central Honshu, Japan Reviewed

Tani, N., Tomaru, N., Tsumura, Y., Araki, M., and Ohba, K

Journal of Plant Research   Vol. 111 ( 1101 ) page： 7-15   1998

Detection of quantitative trait loci for juvenile growth, flower bearing and rooting ability based on a linkage map in sugi Reviewed

Yoshimaru, H, Ohba, K., Tsurumi, K., Tomaru, N., Murai, M, Mukai, Y., Suyama, Y., Tsumura. Y., Kawahara, T., and Sakamaki, Y.

Theoretical and Applied Genetics   Vol. 97 ( 1-2 ) page： 45-50   1998

Intraspecific variation and phylogeographic patterns of Fagus crenata (Fagaceae) mitochorial DNA Reviewed

Tomaru, N., Takahashi, M., Tsumura, Y., Takahashi, M., and Ohba, K

American Journal of Botany   Vol. 85 ( 5 ) page： 629-636   1998

Genetic diversity in Fagus crenata (Japanese beech): influence of the distributional shift during the late-Quaternary Reviewed

Tomaru, N., Mitsutsuji, T., Takahashi, M., Tsumura, Y., Uchida, K., and Ohba, K.

Heredity   Vol. 78 ( 3 ) page： 241-251   1997

Linkage analysis of isozyme and dwarf loci, and detection of lethal genes in sugi (Cryptomeria japonica D. Don) Reviewed

Kuramoto, N., Tomaru, N., Murai, M., and Ohba, K.

Breeding Science   Vol. 47 ( 3 ) page： 259-266   1997

Allozyme variation in natural populations of hinoki, Chamaecyparis obtusa and its comparison with hinoki plus-trees selected from artificial stands Reviewed

Uchida, K., Tomaru, N., Tomaru, T., Yamamoto, C., and Ohba, K.

Breeding Science   Vol. 47 ( 1 ) page： 7-14   1997

Regeneration system and genetic diversity of Cryptomeria japonica at different growing altitudes Reviewed

Taira, H., Tsumura, Y., Tomaru, N., and Ohba, K.

Canadian Journal of Forest Research   Vol. 27 ( 4 ) page： 447-452   1997

Genetic diversity and differentiation in populations of Japanese stone pine (Pinus pumila) in Japan Reviewed

Tani, N., Tomaru, N., Araki, M., and Ohba, K.

Canadian Journal of Forest Research   Vol. 26 ( 8 ) page： 1454-1462   1996

Genetic analysis of four isozyme loci and a linkage relationship between dwarf gene (dw) and leaf whiting gene in summer (swl) of sugi (Cryptomeria japonica) Reviewed

Kuramoto, N., Tomaru, N., Murai, M., Ohba, K.

Breeding Science   Vol. 46 ( 4 ) page： 379-384   1996

The genetic variation of isozyme in natural populations of masson pine. Reviewed

Huang, Q., Wang, L., Tomaru, N., and Ohba, K.

Acta Genetica Sinica   Vol. 22 ( 2 ) page： 142-151   1995

Molecular phylogeny of conifers using RFLP analysis of PCR-amplified specific chloroplast genes Reviewed

Tsumura, Y., Yoshimura, K., Tomaru, N., and Ohba, K.

Theoretical and Applied Genetics   Vol. 91 ( 8 ) page： 1222-1236   1995

Allozyme variations and spatial distributions of genotypes in strains of a butt-rot fungus of Japanese larch Reviewed

Kuroda, Y., Tomaru, N., and Hanyu, M.

Journal of the Japanese Forestry Society   Vol. 77 ( 5 ) page： 480-485   1995

A linkage map for sugi(Cryptomeria japonica)based on RFLP, RAPD and isozyme loci Reviewed

Mukai, Y., Suyama, Y., Tsumura, Y., Kawahara, T., Yoshimaru, H., Kondo, T., Tomaru, N., Kuramoto, N., and Murai, M.

Theoretical and Applied Genetics   Vol. 90 ( 6 ) page： 835-840   1995

Correlations between the growth vigour and isozyme phenotypes of Pinus massoniana Lamb. Reviewed

Huang, Q., Wang, L., Tomaru, N. and Ohba, K.

Scientia Silvae Sinicae   Vol. 30 ( 3 ) page： 214-219   1994

Genetic variation and population differentiation in natural populations of Cryptomeria japonica Reviewed

Tomaru, N., Tsumura, Y., and Ohba, K.

Plant Species Biology   Vol. 9 ( 3 ) page： 191-199   1994

Genetic control of isozyme variation in masson pine, Pinus massoniana Lamb Reviewed

Huang, Q.-Q., Tomaru. N., Wang, L.-H., and Ohba, K.

Silvae Genetica   Vol. 43 ( 5-6 ) page： 285-292   1994

Studies on genetic variation in plantations and plus-trees of Cryptomeria Japonica using isozyme markers Reviewed

Tomaru, N.

Memoirs of The Institue of Agriculture and Forestry, University of Tsukuba   Vol. 6   page： 1-66   1994

Allozyme variation in plus-tree of hinoki, Chamaecyparis obtusa, selected from artificial stands Reviewed

Uchida, K., Tomaru, N., Tsumura, Y., Takahashi, C. and Ohba, K.

Japanese Journal of Breeding   Vol. 43 ( 4 ) page： 485-494   1993

Allozyme variation in artificial stands and a plus-tree group of sugi, Cryptomeria japonica in Ibaraki Prefecture Reviewed

Tomaru, N., Tsumura, Y. and Ohba, K.

Journal of the Japanese Forestry Society   Vol. 74 ( 1 ) page： 44-48   1992

Inheritance of isozyme variants in Korean pine(Pinus koraiensis) Reviewed

Tomaru, N., Tsumura, Y. and Ohba, K.

Journal of the Japanese Forestry Society   Vol. 72 ( 3 ) page： 194-200   1990

種内の遺伝的変異−中立変異−. 井出雄二・白石進編 森林遺伝育種学

戸丸信弘, 津村義彦（ Role： Contributor）

文永堂出版  2012 

Conservation of Biological and Genetic Resources by Conservation Genetic Approaches (Considering Biodiversity as Resources. Imura, H. ed.) (in Japanese)

Nobuhiro Tomaru（ Role： Contributor）

Cokusai Shoin Co., Ltd.  2010 

Genetic Variation and Structure in Fagus crenata populations (Applied Ecology for Restoration for Beech Forests. Terazawa, K. and Koyama, H. eds.) (in Japanese)

Nobuhiro Tomaru（ Role： Contributor）

Bun-ichi Sogo Shuppan Co.  2008 

Encyclopedia of Plant Breeding (Japanese Society of Breeding ed.) (in Japanese)

Nobuhiro Tomaru（ Role： Contributor）

Baifukan Co., Ltd.  2005 

Forest management and conservation using microsatellite markers: the example of Fagus (Biotechnology in Agriculture and Forestry, Vol. 55 Molecular Marker Systems. Lorz, H. L. and Wenzel, G. eds.)

Tsumura, Y., Takahashi, M., Takahashi, T., Tani, N., Asuka Y. and Tomaru N.（ Role： Contributor）

Springer-Verlag  2004 

Fagus crenata, Cryptomeria japonica, and Chamaecyparis obtusa (Tree Ecophysiology Koike, T. ed.) (in Japanese)

Nobuhiro Tomaru（ Role： Contributor）

Asakura Publishing Co., Ltd.  2004 

Encyclopedia of Ecology (Iwasa, Y., Matsumoto, T., Kikuzawa, K., and Ecological Society of Japan eds.) (in Japanese)

Nobuhiro Tomaru（ Role： Contributor）

Kyoritsu Shuppan, Co., Ltd.  2003 

History of genes: History and Genetic Variation of Fagus crenata populations (Molecular Ecology of Woody Species. The Society for the Study of Species Biology ed.) (in Japanese)

Nobuhiro Tomaru（ Role： Contributor）

Bun-ichi Sogo Shuppan Co.  2001 

Assessing future climate risks for 11 major tree species using spatial modeling of adaptive genetic variations

Grant number：21H02240  2021.4 - 2024.3

Grants-in-Aid for Scientific Research  Grant-in-Aid for Scientific Research (B)

Uchiyama Kentaro

Elucidation of local environmental adaptation genes of trees using large-scale provenance test forests

Grant number：21H04732  2021.4 - 2024.3

Grants-in-Aid for Scientific Research  Grant-in-Aid for Scientific Research (A)

Tsumura Yoshihiko

Elucidation of adaptive and neutral evolution processes in Fagus crenata using population genomics

Grant number：20H03027  2020.4 - 2023.3

Grants-in-Aid for Scientific Research  Grant-in-Aid for Scientific Research (B)

Tomaru Nobuhiro

Effects of a domestic exotic species, Magnolia kobus, on the persistence of a rare species, M. stellata

Grant number：17H03830  2017.4 - 2020.3

Grants-in-Aid for Scientific Research  Grant-in-Aid for Scientific Research (B)

Tomaru Nobuhiro

ブナ林の断片化がブナ集団の遺伝的多様性と繁殖に及ぼす影響

2013.4 - 2016.3

科学研究費補助金  基盤研究(B)

戸丸信弘

ブナ集団における適応的遺伝子の探索と生態的意義

2008.4 - 2011.3

科学研究費補助金  基盤研究(B)

戸丸　信弘

針葉樹の雑種苗の分子識別と起源推定

2007.4 - 2010.3

科学研究費補助金  基盤研究(B)

津村義彦

葉緑体ゲノムのSNPを用いたブナの分子系統地理学的・環境適応的研究

2007.4 - 2010.3

科学研究費補助金  基盤研究（B)

高橋誠

フタバガキ科の系統地理学的研究と産地識別のための塩基配列データベースの構築

2006.4 - 2010.3

科学研究費補助金  基盤研究（A)（海外学術調査）

津村義彦

湿地林を構成する希少木本種の繁殖と更新に及ぼす遺伝的荷重の影響の解明

2006.4 - 2009.3

科学研究費補助金  基盤研究（B)

石田清

Studies on gene flow and inbreeding depression in Magnolia stellata for its conservation

2004.4 - 2006.3

Grants-in-Aid for Scientific Research  Grant-in-Aid for Scientific Research (B)

TOMARU Nobuhiro

Characterizations of Amazonian Woods in Relation to the Evolution of Woody Plants

2003.4 - 2006.3

Grants-in-Aid for Scientific Research  Grant-in-Aid for Scientific Research (A)

FUKUSHIMA Kazuhiko, YAMAMOTO Hiroyuki, TOMARU Nobuhiro, MATSUSHITA Yasuyuki, SUZUKI Mitsuo, MARUYAMA Tsuyoshi

森林生態系の維持機構に関する個体群統計学・分子集団遺伝学的研究

2003.4 - 2004.3

科学研究費補助金  基盤研究（C)(1)（企画調査）

山本進一

分子生態遺伝学的手法による森林動態に関する研究

2002.4 - 2005.3

科学研究費補助金  基盤研究(A)

山本進一

メソスケールでの森林現象動態モニタリング技術の基盤確立

2001.4 - 2004.3

科学研究費補助金  基盤研究(A)(2)(展開)

山本進一

日本海側ブナ林構成樹種における地理的分布と遺伝的構造の歴史的関連性

2000.4 - 2003.3

科学研究費補助金  基盤研究(C)(2)(一般)

戸丸信弘

マイクロサテライトマーカーによる照葉樹天然林の繁殖構造解析

1999.4 - 2002.3

科学研究費補助金  基盤研究(B)(2)(一般)

山本進一

オルガネラDNA解析によるブナ集団の系統地理学的研究

1998.4 - 2000.3

科学研究費補助金  基盤研究（C)

戸丸信弘

Biostatistics Exercise

2024

生物環境科学実験実習2

2024

生物環境科学実験実習1

2024

生物学実験

2024

生命農学序説

2024

Fundamentals of Biology I

2023

Forest Biology

2023

Conservation Genetics

2023

Basic Practical Courses

2023

Forest Resource Management 1

2023

生物環境科学実験実習１

2023

生命農学序説

2023

Conservation Genetics

2022

Plant Taxonomy and Vegetation Science

2022

Fundamentals of Biology I

2022

Basic Practical Courses

2022

Forest Biology

2022

Forest Resource Management 1

2022

生命農学序説

2022

生物環境科学実験実習１

2022

生物環境科学実験実習２

2022

環境問題と人間

2021

Basic Practical Courses

2021

Forest Biology

2021

Laboratory and Practice in Bioenvironmental Sciences 1

2021

Plant Taxonomy and Vegetation Science

2021

Conservation Genetics

2021

Laboratory and Practice in Bioenvironmental Sciences 2

2021

Forest Resource Management 1

2021

Forest Resource Management 1

2020

Conservation Genetics

2020

Forest Biology

2020

Laboratory/Field Course in Forest Biology

2020

Laboratory and Practice in Bioenvironmental Sciences 2

2020

Laboratory and Practice in Bioenvironmental Sciences 1

2020

Environmental Problems and Human Beings

2020