Updated on 2021/04/12

写真a

 
FUJIWARA Shin-ichi
 
Organization
Nagoya University Museum Lecturer
Graduate School
Graduate School of Environmental Studies
Title
Lecturer

Degree 3

  1. Doctor (Science) ( 2008.3   The University of Tokyo ) 

  2. 修士(理学) ( 2005.3   東京大学 ) 

  3. 学士(理学) ( 2003.3   東京大学 ) 

Research Interests 4

  1. vertebrate paleontology

  2. Functional morphology

  3. Functional Morphology

  4. Vertebrate Paleontology

Research Areas 5

  1. Others / Others  / Comparative Anatomy

  2. Others / Others  / Vertebrate Paleontology

  3. Others / Others  / Functional Morphology

  4. Natural Science / Biogeosciences  / Functional Morphology

  5. Natural Science / Biogeosciences  / Vertebrate Paleontology

Current Research Project and SDGs 2

  1. 四肢動物の肩帯の運動機能適応の進化

  2. 四肢動物の骨格形態と四足歩行様式の関係の理解、および絶滅動物の古生態復元への応用

Research History 4

  1. 東京大学総合研究博物館・特任助教   特任助教

    2012.4 - 2012.10

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    Country:Japan

  2. 東京大学総合研究博物館・日本学術振興会特別研究員PD   研究員

    2010.4 - 2012.3

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    Country:Japan

  3. 東京大学総合研究博物館・特任研究員   研究員

    2009.10 - 2010.3

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    Country:Japan

  4. 東京大学総合研究博物館・特任研究員   研究員

    2009.4 - 2009.6

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    Country:Japan

Education 3

  1. The University of Tokyo   Graduate School, Division of Science

    2005.4 - 2008.3

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    Country: Japan

  2. The University of Tokyo   Graduate School, Division of Science

    2003.4 - 2005.3

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    Country: Japan

  3. The University of Tokyo   Faculty of Science

    1999.4 - 2003.3

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    Country: Japan

Professional Memberships 3

  1. Society of Vertebrate Paleontology

  2. Palaeontological Society of Japan

  3. 日本哺乳類学会

Awards 1

  1. 学術賞

    2018.6   日本古生物学会   脊椎動物の機能形態学的研究

 

Papers 22

  1. The range of atlanto-occipital joint motion in cetaceans reflects their feeding behavior. Reviewed International journal

    Taro Okamura, Shin-Ichi Fujiwara

    Journal of Anatomy   Vol. 236 ( 3 ) page: 434 - 447   2020.3

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    Authorship:Last author   Publishing type:Research paper (scientific journal)   Publisher:WILEY  

    The cetaceans display a wide variety of lifestyles, especially with regard to their feeding behavior. However, the evolutionary process of the feeding behavior in cetaceans is still poorly understood, in part because reconstructing the feeding behavior of extinct taxa remains difficult. In cetaceans, cranium mobility relative to the trunk largely depends on the range of motion permitted by the atlanto-occipital joint, given the lack of flexibility of the cervical series. In this study, we examined 56 extant cetacean skeletal specimens from 30 species in 25 genera and nine families in order to investigate the relationships between anatomical traits and feeding behavior. Our results suggest that the range of dorso-ventral motion allowed by the atlanto-occipital joint (ROM) depends on prey habitat and the feeding technique of cetaceans. Cetaceans feeding on benthic/demersal prey had a relatively large ROM compared with those feeding on pelagic prey. In addition, ROM was largest in raptorial feeders, intermediate in suction feeders, and smallest in ram-filter feeders. Among raptorial feeders, ROM tended to be larger in taxa that facultatively tear off the prey's flesh compared with taxa that swallow their prey whole. Therefore, we conclude that ROM is a powerful tool to reliably reconstruct the feeding behavior of extinct cetacean taxa.

    DOI: 10.1111/joa.13111

    Web of Science

    PubMed

  2. The sandwich structure of keratinous layers controls the form and growth orientation of chicken rhinotheca. Reviewed International journal

    Yukine Urano, Yasunobu Sugimoto, Kyo Tanoue, Ryoko Matsumoto, Soichiro Kawabe, Tomoyuki Ohashi, Shin-Ichi Fujiwara

    Journal of Anatomy   Vol. 235 ( 2 ) page: 299 - 312   2019.8

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    Authorship:Last author   Publishing type:Research paper (scientific journal)   Publisher:WILEY  

    The upper beak bone of birds is known to be overlain by the rhinotheca, which is composed of the horny sheath of keratinous layers. However, the details of the structure and growth pattern of the rhinotheca are yet to be understood. In this study, the microstructure of the rhinotheca from chicken specimens of different growth stages (ranging from 1 to ~ 80 days old) was analyzed using a combination of thin section and scanning electron microscopy observations, and small-angle X-ray scattering analysis. We found that the rhinotheca comprises three different layers - outer, intermediate, and inner layers - throughout its growth. The outer layer arises from the proximal portion of the beak bone and covers the dorsal surface of the rhinotheca, whereas the intermediate and inner layers originate in the distal portion of the beak bone and underlie the outer layer. This tri-layered structure of the rhinotheca was also observed in wild bird specimens (grey wagtail, king quail, and brown dipper). On the median plane, micro-layers making up the outer and inner layers are bedded nearly parallel to the rostral bone at the base. However, more distally positioned micro-layers of the outer layer are more anteverted distally. The micro-layers of the intermediate layer are bedded nearly perpendicular to those of the outer and inner layers on the median plane. The growth of micro-layers in the intermediate layer adds thickness to the rhinotheca, which causes the difference in profile between the beak bone and the rhinotheca in the distal portion of the beak. Moreover, the entire intermediate layer grows distally as new proximal micro-layers form. The outer layer is dragged distally by the intermediate layer as a result of its distal growth, for the three layers are closely packed to each other at their boundaries. Furthermore, the occurrence of the intermediate and inner layers in the distal portion of the rostral bone may be because the distal end of the beak is frequently used and worn, and the rhinotheca therefore needs to be replaced more frequently at the distal end. The rhinotheca structure described here will be an important and useful factor in the reconstruction of the beaks of birds in extinct taxa.

    DOI: 10.1111/joa.12998

    Web of Science

    PubMed

  3. Feeding Behavior and Neck Mobility in the Extinct Long-Snouted Choristodera Champsosaurus (Reptilia: Diapsida) vs Crocodilia Gavialis

    Matsumoto R., Fujiwara S., Evans S. E.

    JOURNAL OF MORPHOLOGY   Vol. 280   page: S176 - S176   2019.6

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  4. The Tri-layered Keratinous Sheath on the Upper Beak Bone of Chicken and its Relevance to the Form and Growth Orientation of the Beak

    Urano Y., Sugimoto Y., Tanoue K., Matsumoto R., Kawabe S., Ohashi T., Fujiwara S.

    JOURNAL OF MORPHOLOGY   Vol. 280   page: S231 - S232   2019.6

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  5. Propping on Forelimbs Carries a High Risk of Rib Fractures in Tyrannosaurus (Dinosauria: Theropoda)

    Fujiwara S., Ando K.

    JOURNAL OF MORPHOLOGY   Vol. 280   page: S123 - S123   2019.6

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  6. Physical constraints on sand crab burrows: Mechanical properties of wet sand explain the size and spatial distributions of burrows on beaches Reviewed

    Ayuko Shinoda, Shin-ichi Fujiwara, Hirofumi Niiya, Hiroaki Katsuragi

    PLoS ONE   Vol. 14 ( 5 )   2019.5

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    Publishing type:Research paper (scientific journal)   Publisher:PUBLIC LIBRARY SCIENCE  

    The diameter and vertical depth of sand crab tunnels in sandy beaches are usually restricted to a few centimeters scale and several tens of centimeters, respectively. We designed a study to determine what physical factors restrict tunnel diameter and predict the maximum attainable tunnel diameter and depth. We collected field data on the size and spatial distributions of ghost crab (Ocypode spp.) burrows on two sandy beaches (Kawage Beach in Tsu, Mie Prefecture, Japan and Sakieda Beach in Ishigaki, Okinawa Prefecture, Japan), where O. ceratophthalma dominants the ghost crab fauna. We measured burrow depths and distance from shoreline in concert with water content of sandy beaches. To explain our observed distributions of crab burrows in the field, we performed experiments in a lab microcosm, comprising a horizontal tunnel through wet sand. We measured the static stability of tunnel structures in relation to water content and two strengths computed from loading force exerted on the sand overlying the tunnels. By comparing field and experimental data, we found that crabs construct their burrows in appropriately wet zones (wet enough to provide sufficient cohesion of the sand grains in tunnel walls to prevent collapse) and that tunnel diameters and depths are sufficiently small to prevent deformation and collapse of their tunnels.

    DOI: 10.1371/journal.pone.0215743

    Web of Science

  7. Experimental Study of Stability and Strength of a Horizontal Tunnel in Wet Granular Layer: Empirical Forms of Strength to Discuss the Mechanical Stability of Crab Burrows

    Shinoda Ayuko, Fujiwara Shin-ichi, Niiya Hirofumi, Katsuragi Hiroaki

    Journal of the Society of Powder Technology, Japan   Vol. 56 ( 4 ) page: 194 - 202   2019

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    Publisher:The Society of Powder Technology, Japan  

    <p>Empirical forms for the strengths characterizing stability of a horizontal tunnel in wet granular matter are experimentally derived. The motivation of this study comes from the observation of crab burrows on sandy beach. To understand the mechanical constraints for constructing the stable burrow structures, we perform a set of simple experiments with a horizontal tunnel structure made in a wet granular layer. Then, the tunnel is compressed from the top of the layer using a universal testing machine. During the compression, the shrinking tunnel shape is captured by a camera. Using the measured compression force, stroke, and shrinking tunnel images, we define and measure the mechanical strengths of wet granular matter with a horizontal tunnel. By systematically varying the experimental conditions, we obtained the empirical forms for estimating strength values. Using the measured results, we briefly discuss the mechanical conditions of sandy crab burrows.</p>

    DOI: 10.4164/sptj.56.194

    CiNii Books

  8. Void structure stability in wet granular matter and its application to crab burrows and cometary pits. Reviewed International journal

    Ayuko Shinoda, Shin-Ichi Fujiwara, Hirofumi Niiya, Hiroaki Katsuragi

    Scientific Reports   Vol. 8 ( 1 ) page: 15784 - 15784   2018.10

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    Language:English   Publishing type:Research paper (scientific journal)   Publisher:NATURE PUBLISHING GROUP  

    Cohesive granular matter can support stable void structures, which can universally be found in various scenes from everyday lives to space. To quantitatively characterize the stability and strength of a void structure in cohesive granular matter, we perform a simple tunnel-compression experiment with wet granular matter. In the experiment, a horizontal tunnel in a wet granular layer is vertically compressed with a slow compression rate. The experimental result suggests that the tunnel deformation can be classified into the following three types: (i) shrink, (ii) shrink with collapse, and (iii) subsidence by collapse. Using the experimental result, we estimate the stable limit of various void structures in a cohesive granular layer from crab burrows on a sandy beach to the pits observed on cometary surfaces.

    DOI: 10.1038/s41598-018-33978-8

    Web of Science

    PubMed

  9. Fitting unanchored puzzle pieces in the skeleton: appropriate 3D scapular positions for the quadrupedal support in tetrapods. Reviewed International journal

    Shin-Ichi Fujiwara

    Journal of Anatomy   Vol. 232 ( 5 ) page: 857 - 869   2018.5

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    Authorship:Lead author   Language:English   Publishing type:Research paper (scientific journal)   Publisher:WILEY  

    Deducing the scapular positions of extinct tetrapod skeletons remains difficult, because the scapulae and rib cage are connected with each other not directly by skeletal joint, but by thoracic muscles. In extant non-testudine quadrupedal tetrapods, the top positions of the scapulae/suprascapulae occur at the anterior portion of the rib cage, above the vertebral column and near the median plane. The adequacy of this position was tested using three-dimensional mechanical models of Felis, Rattus and Chamaeleo that assumed stances on a forelimb on a single side and the hindlimbs. The net moment about the acetabulum generated by the gravity force and the contractive forces of the anti-gravity thoracic muscles, and the resistance of the rib to vertical compression between the downward gravity and upward lifting force from the anti-gravity thoracic muscle depend on the scapular position. The scapular position common among quadrupeds corresponds to the place at which the roll and yaw moments of the uplifted portion of the body are negligible, where the pitch moment is large enough to lift the body, and above the ribs having high strength against vertical compression. These relationships between scapular position and rib cage morphology should allow reliable reconstruction of limb postures of extinct taxa.

    DOI: 10.1111/joa.12778

    Web of Science

    PubMed

  10. How does the curvature of the upper beak bone reflect the overlying rhinotheca morphology? Reviewed International journal

    Yukine Urano, Kyo Tanoue, Ryoko Matsumoto, Soichiro Kawabe, Tomoyuki Ohashi, Shin-Ichi Fujiwara

    Journal of Morphology   Vol. 279 ( 5 ) page: 636 - 647   2018.5

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    The beak has independently been evolved accompanied by the edentulism in many tetrapod linages, including extant Testudinata and Aves, and its form and function have been greatly diversified. The beak is formed by beak bones and the overlying keratinous cover, although their profiles are different from each other. Therefore, it is difficult to reliably reconstruct the entire profile of the beak in extinct taxa, whose keratinous tissues are rarely preserved. For elucidation of the morphological relationship between beak bone and overlying keratinous cover, we compared the curvature distribution of the culminal profiles of the upper beak bone and the overlying keratinous cover (rhinotheca) with each other using CT-scan, in 66 extant testudinatan and avian specimens (Aves: 33 genera, 24 families; Testudinata: 12 genera seven families). In both, rhinotheca and beak bone, the curvature of the profile was nearly constant rostral to a certain point, which was defined as the transition point, and the transition points of the rhinotheca and beak bone were close to each other. The profiles of the rhinotheca and beak bone rostral to their transition point were different in curvature and length. However, the ratio between the curvatures of rhinotheca and the beak bone strongly correlated with the arc angle of the rostral culminal profiles of the beak bone. The upper beak profile in extinct taxa is expected to be reconstructed more reliably using the abovementioned relationship between the beak bone and the rhinotheca.

    DOI: 10.1002/jmor.20799

    Web of Science

    PubMed

  11. A new embedding method for preparing thin section of animal hard and soft tissues

    Urano Yukine, Takagi Natsuko, Tanoue Kyo, Fujiwara Shin-ichi

    Fossils   Vol. 103 ( 0 ) page: 51 - 59   2018

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    Publisher:Palaeontological Society of Japan  

    We propose a new method of embedding animal samples in epoxy resin Devcon ET (Illinois Tool Works Inc.). This new method provides simultaneous observations of hard and soft tissues of extant animals on thin sections. Comparing to methods previously proposed, our method has advantage that the resin is inexpensive, colorless and transparent. The procedure is; (1) dehydrate the formalin-fixed samples sufficiently with acetone; (2) impregnate the sample with a heated mixed solution of Devcon ET and 99.8 % acetone under reduced pressure; (3) impregnate the sample with Devcon ET under reduced pressure and heat it until the resin cures. The second procedure enables to lower the viscosity of the resin. By this method, it became possible to make colorless transparent biological thin sections at low cost. The sample should more effectively be impregnated by the resin with a longer period of the dehydration and drilling of small holes on the surface.

    DOI: 10.14825/kaseki.103.0_51

    CiNii Books

  12. Farewell to life on land-thoracic strength as a new indicator to determine paleoecology in secondary aquatic mammals Reviewed

    ANDO Konami, FUJIWARA Shin-ichi

    Journal of Anatomy     2016

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    Language:English   Publishing type:Research paper (scientific journal)  

    DOI: 10.1111/joa.12518

  13. Crabs grab strongly depending on mechanical advantages of pinching and disarticulation of chela

    FUJIWARA Shin-ichi, KAWAI Hiroki

    Journal of Morphology   Vol. 277   page: 1259-1272   2016

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    Authorship:Lead author   Language:English   Publishing type:Research paper (scientific journal)  

    DOI: 10.1002/jmor.20573

  14. Comparative morphological examinations of the cervical and thoracic vertebrae and related spinal nerves in the two-toed sloth Reviewed

    Endo H, Hashimoto O, Taru H, Sugimura K, Fujiwara S-I, Itou T, Koie H, Kitagawa M, & Sakai T.

    Mammal Study   Vol. 38   page: 217-224   2013

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    Language:English   Publishing type:Research paper (scientific journal)  

  15. Elbow joint adductor moment arm as an indicator of forelimb posture in extinct quadrupedal tetrapods Reviewed

    Shin-ichi Fujiwara and John R. Hutchinson

    Proceedings of the Royal Society B: Biological Sciences   Vol. 279   page: 2561-2570   2012.2

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    Authorship:Lead author   Language:English   Publishing type:Research paper (scientific journal)  

    DOI: 10.1098/rspb.2012.0190

  16. Topsy-turvy locomotion: biomechanical specializations of the elbow in suspended quadrupeds reflect inverted gravitational constraints

    Shin-ichi Fujiwara, Hideki Endo, and John R. Hutchinson

    Journal of Anatomy   Vol. 219 ( 2 ) page: 176-191   2011.4

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    Authorship:Lead author   Language:English   Publishing type:Research paper (scientific journal)  

  17. A sub-adult growth stage indicated in the degree of suture co-ossification in Triceratops. Reviewed

    Shin-ichi Fujiwara and Yuji Takakuwa

    Bulletin of Gunma Museum of Natural History   Vol. 15   page: 1-17   2011

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    Authorship:Lead author   Language:English   Publishing type:Research paper (scientific journal)  

  18. Shape of articular surface of crocodilian (Archosauria) elbow joints and its relevance to sauropsids Reviewed

    Shin-ichi Fujiwara, Hajime Taru, and Daisuke Suzuki

    Journal of Morphology   Vol. 271 ( 7 ) page: 883-896   2010.4

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    Authorship:Lead author   Language:English   Publishing type:Research paper (scientific journal)  

    DOI: 10.1002/jmor.10846

  19. A reevaluation of the manus structure in Triceratops (Ceratopsia: Ceratopsidae) Reviewed

    Fujiwara, S.-I.

    Journal of Vertebrate Paleontology   Vol. 29   page: 1136-1147   2009.12

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    Language:English   Publishing type:Research paper (scientific journal)  

    DOI: 10.1671/039.029.0406

  20. Relationship between scapular position and structural strength of rib cage in quadruped animals Reviewed

    Shin-ichi Fujiwara, Osamu Kuwazuru, Norihisa Inuzuka, Nobuhiro Yoshikawa

    Journal of Morphology   Vol. 270 ( 9 ) page: 1084-1094   2009.4

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    Authorship:Lead author   Language:English   Publishing type:Research paper (scientific journal)  

    DOI: 10.1002/jmor.10744

  21. Olecranon orientation as an indicator of elbow joint angle in the stance phase, and estimation of forelimb posture in extinct quadruped animals Reviewed

    Shin-ichi Fujiwara

    Journal of Morphology   Vol. 270 ( 9 ) page: 1107-1121   2009.4

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    Language:English   Publishing type:Research paper (scientific journal)  

    DOI: 10.1002/jmor.10748

  22. Relay strategy and adaptation to a muddy environment in Isselicrinus (Isselicrinidae: Crinoidea). Reviewed

    Fujiwara, S.-I., T. Oji, Y. Tanaka, and Y. Kondo

    Palaios   Vol. 20   page: 241-248   2005.6

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    Language:English   Publishing type:Research paper (scientific journal)  

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Books 5

  1. ifの地球生命史 : "もしも"絶滅した生物が進化し続けたなら

    土屋 健(サイエンスライター), 服部 雅人, 藤原 慎一, 椎野 勇太

    技術評論社  2021  ( ISBN:9784297119201

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  2. 語源が分かる恐竜学名辞典 : 恐竜類以外の古生物(翼竜類・魚竜類など)の学名も一部含む

    松田, 眞由美, 小林, 快次, 藤原, 慎一

    北隆館  2017.1  ( ISBN:9784832607347

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    Total pages:540p   Language:Japanese

    CiNii Books

  3. 見る目が変わる博物館の楽しみ方 地球・生物・人類を知る

    三河内彰子、三河内岳、椎野勇太、実吉玄貴、藤原慎一、折原貴道、矢野興一、鵜沢美穂子、皆木宏明、黒木真理、松本涼子、相川稔、松原始、佐野勝宏、鶴見英成、石井龍太( Role: Sole author)

    ペレ出版  2016.10  ( ISBN:978-4-86064-490-1

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    Total pages:435   Language:Japanese Book type:General book, introductory book for general audience

  4. 恐竜学入門―かたち・生態・絶滅―

    真鍋真、藤原慎一、松本涼子( Role: Joint translator)

    東京化学同人  2015.1  ( ISBN:978-4-8079-0856-1

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    Total pages:396   Language:Japanese Book type:Textbook, survey, introduction

  5. 恐竜戦国時代の覇者!トリケラトプス~知られざる大陸ララミディアでの攻防~

    藤原慎一・塚腰実・林昭次( Role: Joint editor)

    読売新聞社  2014 

MISC 3

  1. The record of 25th Nagoya University Museum special exhibition : "Nagoya University Campus Museum" for the 80th Anniversary since its foundation

    UMEMURA Ayako, UJIHARA Kimiko, NISHIDA Sachiko, KADOWAKI Seiji, FUJIWARA Shin-ichi

      ( 35 ) page: 55 - 68   2020

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  2. Preparing PEG-impregnated crustacean specimens that retain the joint flexibilities

    FUJIWARA Shin-ichi

      ( 32 ) page: 27 - 32   2017

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  3. The mechanical property of a tunnel structure in wet granular layer

    Shinoda Ayuko, Fujiwara Shin-ichi, Katsuragi Hiroaki

    Meeting Abstracts of the Physical Society of Japan   Vol. 72 ( 0 ) page: 2901 - 2901   2017

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    Language:Japanese   Publisher:The Physical Society of Japan  

    DOI: 10.11316/jpsgaiyo.72.1.0_2901

Research Project for Joint Research, Competitive Funding, etc. 3

  1. 絶滅動物の胴体の形状と重心位置に基づく体肢姿勢の復元:四足歩行と二足歩行の姿勢進化について

    2017.4 - 2018.3

    公益財団法人藤原ナチュラルヒストリー振興財団学術(動物学)研究助成 

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    Grant type:Competitive

  2. 鍾乳洞の正体をあばこう!

    2016.4 - 2017.3

    日本学術振興会 ひらめき★ときめきサイエンス 

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    Grant type:Competitive

  3. Mechanical model of the elbow joint angles in the animals without prominent olecranons

    2009.6 - 2009.10

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    Grant type:Competitive

KAKENHI (Grants-in-Aid for Scientific Research) 13

  1. 比較解剖学的・組織学的手法による古生物の嘴の復元

    2018.4 - 2022.3

    科学研究費補助金  基盤研究(C)

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  2. 比較解剖学的・組織学的手法による古生物の嘴の復元

    Grant number:18K03828  2018.4 - 2022.3

    日本学術振興会  科学研究費助成事業 基盤研究(C)  基盤研究(C)

    田上 響

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    骨と角質からなり、さまざまな形態と機能を持つ脊椎動物のクチバシを、化石分類群でより確からしく復元することを目指し、平成30年度に引き続き令和元年度に、現生鳥類標本を用いて角質部の微細構造を解析した。さらに紫外線蛍光撮影により、現生および化石鳥類の頭骨標本を対象とした角質部の分布調査を行った。
    まず現生鳥類のクチバシを用いた角質部の微細構造の解析を、(1) 薄片による角質部断面の偏光顕微鏡観察、(2) 角質部断面の走査型電子顕微鏡観察、(3) X線小角散乱法の3手法により行った。(1)により、クチバシの角質部は外層、中間層、内層の3層に分かれること、また(2)により、外層はさらに微細な薄層の重なりからなること、そして(3)により、層ごとに薄層の傾きが異なることが明らかとなった。
    また紫外線蛍光撮影により、古生物のクチバシの復元を試みた。現生鳥類の複数の分類群で頭骨の紫外線照射下撮影を行ったところ、嘴の角質部に覆われる領域は概ね一つの色を呈し、その後端付近において、色の異なる領域との境界が見られた。また、漸新芦屋層群産鳥類化石の下顎標本を同様に撮影したところ、表面に色の異なる領域が確認された。一つの色を呈する領域が必ずしも左右対称でないことから、標本の保存状態等の解釈を要するものの、この調査によって、古生物のクチバシにおける角質部の分布を、紫外線蛍光撮影により検証できる可能性が示唆された。
    さらに、CT撮像およびフォトグラメトリによる現生および化石標本の三次元データの収集を行い、角質部とクチバシに伸びる三叉神経の分布に関する調査も開始した。
    クチバシの角質部の微細構造解析において、偏光顕微鏡を用いクロスニコルで現生鳥類のクチバシの正中断面の薄片観察を行うと、角質部で消光位が異なる三層(外層、中間層、内層)の層構造を確認できた。また、走査型電子顕微鏡下観察においては、外層内に更に微細な層構造があることが確認でき、この層の向きが偏光顕微鏡下観察での消光位と関係することが分かった。さらにX線小角散乱法によって、外層では背腹軸で一定間隔の繰り返し構造があることと、外層と中間層では微細構造が異なることも示された。この結果を令和元年7月の第12回国際脊椎動物形態学会にて報告した。また、本調査の結果をまとめた論文がJournal of Anatomyにて出版された。
    紫外線蛍光撮影による角質部の分布調査では、まず現生鳥類の頭骨標本の紫外線蛍光撮影を行った。複数の標本で角質部に覆われる領域は概ね一つの色を呈し、その後端付近において、色の異なる領域との境界が見られた。漸新統芦屋層群より産出した、化石鳥類の下顎標本の紫外線蛍光撮影を行った。撮影の結果、表面に色の異なる領域が確認された。左右外側で色の分布域が異なるため、可視光では確認できない角質部の分布を、紫外線蛍光撮影により調査できる可能性が示唆される。
    さらに、現生および化石標本を対象にCT撮像を行い、CTスキャナで撮像できない大きさの化石標本では、フォトグラメトリによる三次元データの収集を行うことで、角質部とクチバシに伸びる三叉神経の分布に関する調査も開始した。三叉神経を調査することにより、クチバシの角質部の分布を間接的に復元できると期待される。
    紫外線蛍光撮影による角質部の分布調査は、鳥類では現生標本と化石標本を比較できる点で、今後も有望であると考えられる。ただ現生カメ類標本はこれまで調査できた数が少なく明瞭な結果も出なかったため、今後撮影を継続し、鳥類と同様の結果を得られるか確認する。クチバシを持っていたと考えられる化石標本に関しても、撮影の許可を頂いたものより紫外線照射下撮影を進め、現生標本と比較しつつ、角質部の分布を検証する予定である。ただ現時点では、紫外線蛍光撮影での色の違いをもたらす要因が明らかとなっていない。今後現生標本を対象に、角質部と骨質部をつなぐ結合組織の分布調査を予定している。
    令和元年度より開始したクチバシの領域に伸びる三叉神経の分布調査により、間接的に角質部の分布を復元できることが期待される。まず現生標本での角質部と三叉神経の分布領域の関係を確認し、化石標本での角質部復元を目指す。現時点で、歯とクチバシをともに持つとされる角竜類恐竜のデータを収集しており、まずこれらの解析から進める予定である。

  3. Evolution of neck motion system in tetrapods, revealing from the morphology of occipital condyles

    Grant number:17K05698  2017.4 - 2022.3

    Japan Society for the Promotion of Science  Grants-in-Aid for Scientific Research Grant-in-Aid for Scientific Research (C)  Grant-in-Aid for Scientific Research (C)

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    Authorship:Coinvestigator(s) 

  4. 後頭部の形態から読み解く四肢動物の首の運動機能の進化

    2017.4 - 2021.3

    科学研究費補助金  基盤研究(C)

  5. Relationship between skeletal morphology and gait system in quadrupedal tetrapods with implications to gait reconstructions of extinct taxa

    Grant number:17K17794  2017.4 - 2021.3

    Japan Society for the Promotion of Science  Grants-in-Aid for Scientific Research Grant-in-Aid for Young Scientists (B)  Grant-in-Aid for Young Scientists (B)

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    Authorship:Principal investigator 

    Grant amount:\4420000 ( Direct Cost: \3400000 、 Indirect Cost:\1020000 )

  6. 濡れた紛体における穴構造の安定性とカニの巣穴強度理解への応用

    2016.4 - 2018.3

    科学研究費補助金 

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    Authorship:Coinvestigator(s) 

  7. Stability of a void structure in wet granular layer and its application to crab burrow strength

    Grant number:16K13861  2016.4 - 2018.3

    Japan Society for the Promotion of Science  Grants-in-Aid for Scientific Research Grant-in-Aid for Challenging Exploratory Research  Grant-in-Aid for Challenging Exploratory Research

    Katsuragi Hiroaki

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    Ghost crabs make cm-scale-diameter burrows on appropriately wet sandy substrate. Physically, such a sandy substrate can be regarded as wet granular matter. Thus, the size and distribution of crab burrows might be governed by the stability of a void structure in wet granular layer. In this study, we performed the experiment and field work to quantitatively reveal the relation among the mechanical properties of tunnel structure in wet granular layer , spatial distribution of crab burrows, and their size distribution. From the obtained results, we found that the actual crab burrows are sufficiently stable in terms of wet granular mechanics.

  8. Function and evolution of beaks in vertebrates

    Grant number:15K05331  2015.4 - 2018.3

    Japan Society for the Promotion of Science  Grants-in-Aid for Scientific Research Grant-in-Aid for Scientific Research (C)  Grant-in-Aid for Scientific Research (C)

    Tanoue Kyo

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    Vertebrate beaks, which are diverse both in morphology and function, comprise bony and keratinous elements. More accurate reconstruction of vertebrate beaks than in previous studies was aimed in this project. Comparative morphology analyses of extant beaked vertebrates indicate keratinous sheaths develop in the direction to prevent growth interruption by bony beaks at least in birds. In addition, observation of thin sections of avian beaks revealed multi-layered structure of the keratinous beaks. The origin of the layer can be deduced from the morphology of the bony beak. Moreover, fine striations are preserved in the rostral region of fossils vertebrates which are interpreted to have been beaked. Based on the distribution of the striations, larger keratinous sheath than in previous reconstruction was presumed in a specimen of ceratopsian dinosaur.

  9. 脊椎動物における嘴の機能と進化

    2015.4 - 2017.3

    科学研究費補助金  基盤研究(C)

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    Authorship:Coinvestigator(s) 

  10. Relationship between scapular morphology and locomotor behavior in tetrapods

    2013.4

    Grant-in-Aid for Scientific Research  Grant-in-Aid for Young Scientists(B)

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    Authorship:Principal investigator 

  11. 鳥類とモグラ類の胸郭の運動適応と進化

    2012.9

    科学研究費補助金 

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    Authorship:Principal investigator 

  12. 四肢動物の前肢骨格形態と運動機能との関係、及び絶滅動物の前肢姿勢復元

    2010.4 - 2012.3

    科学研究費補助金 

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    Authorship:Principal investigator 

  13. 鳥盤類恐竜の前肢及び胸郭に見られる二次的四足歩行の進化と適応

    2006.4 - 2008.3

    科学研究費補助金 

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    Authorship:Principal investigator 

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Teaching Experience (On-campus) 1

  1. 動植物の科学

    2020

Teaching Experience (Off-campus) 6

  1. 生物形態学

    2021 Gifu University)

  2. 機能形態学(絶滅動物の古生態復元)

    2017 Kyoto University)

  3. 恐竜学

    2014 Nihon University)

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    Level:Undergraduate (liberal arts) 

  4. 生命と環境(自然環境と生物)

    2014 - 2019 Nanzan University)

  5. 理科専門基礎英語DI

    2012 Tokyo Gakugei University)

  6. 地質学特別講義III

    2011 University of Tsukuba)

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    Level:Undergraduate (liberal arts) 

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